Selnekovič, Dávid; Jäch, Manfred A.; Kodada, Ján
Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)
Fig. 1. Details of head. A. Elmomorphus brevicornis Sharp, 1888, male head. B. E. paramontanus Kodada, Selnekovič & Jäch sp. nov., labrum, dorsal aspect. C. E. paramontanus, labrum, ventral aspect. D. E. brevicornis, head, ventral aspect. E. E. paramontanus, medio-anterior portion of labrum in ventral aspect with flat setae. F. E. paramontanus, head, ventral aspect. G. E. paramontanus, flat papillae with seta on ventral side of labrum.
Fig. 2. Details of antennae and mouthparts. A. Elmomorphus brevicornis Sharp, 1888, left antenna, dorsal aspect.B. Same, frontal aspect.C. E. brevicornis, lacinia.D. E. brevicornis, right mandible, dorsal aspect. E. E. brevicornis, left mandible, ventral aspect. F. E. paramontanus Kodada, Selnekovič & Jäch sp. nov., lateral sensory area on terminal maxillary palpomere. G. E. paramontanus, maxilla, ventral aspect. H. Same, dorsal aspect.I. E. paramontanus, apical sensory area on terminal maxillary palpomere. J. E. paramontanus, basal sensory area on terminal maxillary palpomere.
Fig. 4. Details of elytra, legs, and abdomen.A. Elmomorphus paramontanus Kodada, Selnekovič & Jäch sp. nov., stridulatory area on anterolateral ventral surface of elytron.B. E. brevicornis Sharp, 1888, dorsal plastron on elytra (scales). C. E. brevicornis, ventral plastron on abdomen (setae). D. E. brevicornis, dorsal face of left mesofemur.E. E. brevicornis, male foreclaws, lateral aspect. F. E. paramontanus, apex of protibia. G. E. brevicornis, male abdomen, ventral aspect. H. E. brevicornis, apex of male ventrite 5. I. E. paramontanus, apex of female ventrite 5. J. E. amamiensis Nomura, 1959, male ventrite 5.
Fig. 3. Details of mouthparts and thorax. A. Elmomorphus brevicornis Sharp, 1888, mentum and submentum. B. E. brevicornis, labium, ventral aspect. C. E. brevicornis, setae in distal portion of ligula. D. E. brevicornis, apical sensory area on terminal labial palpomere. E. E. paramontanus Kodada, Selnekovič & Jäch sp. nov., pronotum. F. E. brevicornis, male head and prosternum. G. E. amamiensis Nomura, 1959, male metaventrite.
Fig. 5. Elmomorphus brevicornis Sharp, 1888 (CKB). A. Hind wing. B. Male sternite VIII. C. Male sternite IX. D. Female sternite VIII. E. Ovipositor, lateral aspect. Scale bars: a =1.0 mm; b= 0.5 mm.
Fig. 72. Elmomorphus curvipes sp. nov., paratype, male from type locality (NMW), TL: 3.20 mm.
Fig. 68. Elmomorphus similis sp. nov., male from China, locality CWBS 354 (CKB), TL: 3.13 mm.
Fig. 70. Elmomorphus dentipes sp. nov., paratype, male from type locality (NMW), TL: 2.90 mm.
Fig. 107. Habitats of selected species of Elmomorphus Sharp, 1888. A. Vietnam, Phong Mỹ, Sông Ô Lâu River (16°31′15.3″ N, 107°15′00.3″ E); E. striatellus Delève, 1968 and E. siamensis Kodada, Selnekovič & Jäch sp. nov. were found in clumps of leaves and on rootlets of Neptunia sp. B. Vietnam, Thành SƠn (20°30′01.4″ N, 105°06′19.3″ E); E. dentipes sp. nov., and E. similis sp. nov. were found in clumps of leaves.C. Vietnam, ThànhSƠn,Kho Mường(20°28′51.3″N,105°07′51.6″E); E. paramontanus Kodada, Selnekovič & Jäch sp. nov., E. similis sp. nov., and E. cf. sulcatus were found on submerged rootlets of bamboo and banana trees. D. Same locality, showing submerged rootlets. E. Vietnam, Trung Mỹ (21°23′35.3″N, 105°42′52.7″E); E. cf. sulcatus was found in clumps of leaves and on submerged rootlets of bamboo, Bauhinia sp., and Sterculia lanceolata Cav. F. Same locality, clump of leaves and partly submerged rootlets.
Fig. 6. Elmomorphus striatellus Delève, 1968, male from Ban Nape, Laos (CKB), TL: 3.50 mm.
Fig. 7. Elmomorphus striatellus Delève, 1968, aedeagus, specimens from Sa Pa, Vietnam (CKB). A. Lateral aspect. B. Lateral aspect. C. Ventral aspect. Scale bar: 0.1 mm.
Fig. 12. Bursa copulatrix and vagina. A. Elmomorphus striatellus Delève, 1968, Cuc Phuong, Vietnam (CKB). B. E. siamensis Kodada, Selnekovič & Jäch sp. nov., paratype, Umphang, Thailand (CKB). C. E. parabrevicornis sp. nov., paratype from type locality (NMW). Scale bar: 0.1 mm.
Fig. 8. Elmomorphus siamensis Kodada, Selnekovič & Jäch sp. nov., paratype, male from Umphang, Thailand (CKB), TL: 3.00 mm.
Fig. 9. Elmomorphus siamensis Kodada, Selnekovič & Jäch sp. nov., paratype from Umphang, Thailand (CKB), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 10. Elmomorphus parabrevicornis sp. nov., holotype (IAECAS), male, TL: 3.12 mm.
Fig. 11. A. Elmomorphus parabrevicornis sp. nov., holotype (IAECAS), aedeagus, lateral aspect. B. Same, ventral aspect. C. E. auripilosus sp. nov., paratype from Vietnam, Tam Dao (CKB), aedeagus, lateral aspect. Scale bar: 0.1 mm.
Fig. 13. Elmomorphus auripilosus sp. nov., paratype, male from Tam Dao, Vietnam (CKB), TL: 3.20 mm.
Fig. 14. Elmomorphus auratus sp. nov., paratype, male from type locality (NMW), TL: 2.90 mm.
Fig. 16. Elmomorphus horaki Kodada, Selnekovič & Jäch sp. nov., paratype, male from type locality (CKB), TL: 2.50 mm.
Fig. 15. Elmomorphus auratus sp. nov., paratype from type locality (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar =0.1 mm.
Fig. 17. Elmomorphus horaki Kodada, Selnekovič & Jäch sp. nov., aedeagus. A. Paratype from type locality (CKB), lateral aspect. B. Same, ventral aspect. C. Paratype from Umphang, Thailand (CKB), lateral aspect. D. Paratype from Umphang, lateral aspect. Scale bar: 0.1 mm.
Fig. 18. Bursa copulatrix and vagina. A. Elmomorphus auripilosus sp. nov., paratype from Cuc Phuong, Vietnam (CKB). B. E. auratus sp. nov., paratype from China, CWBS 246 (CKB). C. E. horaki Kodada, Selnekovič & Jäch sp. nov., paratype from type locality (NMW). Scale bar: 0.1 mm.
Fig. 19. Elmomorphus fusiformis sp. nov., holotype, male (IAECAS), TL: 4.17 mm.
Fig. 21. Elmomorphus elmoides sp. nov., paratype, male from Buon Luoi, Vietnam (CKB), TL: 2.55 mm.
Fig. 22. Elmomorphus elmoides sp. nov., paratype from type locality (CKB), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 28. Bursa copulatrix and vagina. A. Elmomorphus elmoides sp. nov., paratype from type locality (NMW). B. E. calvus sp. nov., paratype from type locality (CKB). C. E. depressus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm.
Fig. 23. Elmomorphus calvus sp. nov., holotype, male (NMW), TL: 3.25 mm.
Fig. 24. Elmomorphus calvus sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 25. Elmomorphus hongkong sp. nov., paratype, male from type locality (NMW), TL: 2.48 mm.
Fig. 26. Elmomorphus hongkong sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 30. Elmomorphus sulcatus sp. nov., holotype (IAECAS), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 31. Elmomorphus corpulentus sp. nov., holotype, male (IAECAS), TL: 3.32 mm.
Fig. 32. Elmomorphus corpulentus sp. nov., holotype (IAECAS), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 33. Elmomorphus ellipticus sp. nov., paratype, male from type locality (NMW), TL: 2.71 mm.
Fig. 34. Elmomorphus ellipticus sp. nov., holotype (IAECAS), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 35. Elmomorphus mazzoldii sp. nov., holotype, male (NMW), TL: 2.63 mm.
Fig. 36. Elmomorphus mazzoldii sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 37. Bursa copulatrix and vagina. A. Elmomorphus ellipticus sp. nov., paratype from type locality (NMW). B. Elmomorphus mazzoldii sp. nov., paratype from type locality (NMW). Scale bar: 0.1 mm.
Fig. 38. Elmomorphus bryanti Hinton, 1935, male from Gombak River, Malaysia (BMNH), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 39. A. Elmomorphus prosternalis Hinton, 1935, holotype (BMNH), prosternal process and median part of metaventrite. B. E. montanus (Grouvelle, 1913), syntype, male (MNHN), prosternal process and median part of metaventrite. C. E. montanus, syntype (MNHN), aedeagus, lateral aspect. D. Same, ventral aspect, detail of parameres and penis. E. Same, ventral aspect. Scale bars: 0.1 mm.
Fig. 41. Elmomorphus paramontanus, Kodada, Selnekovič & Jäch sp. nov., paratype from Thailand (CKB), aedeagus. A. Lateral aspect. B. Ventral aspect, detail of parameres and penis. C. Ventral aspect. Scale bars: 0.1 mm.
Fig. 42. Elmomorphus cuneatus sp. nov., holotype, male (NMW), TL: 2.69 mm.
Fig. 43. Elmomorphus cuneatus sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 44. Elmomorphus umphangicus Kodada, Selnekovič & Jäch sp. nov., paratype, male from type locality (CKB), TL: 2.80 mm.
Fig. 45. Elmomorphus umphangicus Kodada, Selnekovič & Jäch sp. nov., aedeagus. A. Holotype (NMW), lateral aspect. B. Holotype (NMW), ventral aspect. C. Paratype from type locality (CKB), lateral aspect. Scale bar: 0.1 mm.
Fig. 46. Elmomorphus parvulus sp. nov., holotype, male (NMW), TL: 2.54 mm.
Fig. 48. Elmomorphus sausai Kodada, Selnekovič & Jäch sp. nov., holotype, male (NMW), TL: 3.10 mm.
Fig. 52. Elmomorphus longitarsis sp. nov., holotype, male (NMW), TL: 2.93 mm.
Fig. 54. Elmomorphus bispinosus sp. nov., holotype, female (IAECAS), TL: 3.54 mm.
Fig. 55. Bursa copulatrix and vagina. A. Elmomorphus parvulus sp. nov., paratype from type locality (CKB). B. E. yunnanensis Kodada, Selnekovič & Jäch sp. nov., paratype from type locality (CKB). C. E. longitarsis sp. nov., paratype from type locality (NMW). D. E. bispinosus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm.
Fig. 47. Elmomorphus parvulus sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 49. Elmomorphus sausai Kodada, Selnekovič & Jäch sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect, detail of parameres and penis. C. Ventral aspect. Scale bars: 0.1 mm.
Fig. 53. Details of tarsi and aedeagus. A. Elmomorphus yunnanensis Kodada, Selnekovič & Jäch sp. nov., paratype from type locality (NMW), protarsus. B. E. longitarsis sp. nov., paratype fom type locality (NMW), protarsus. C. E. longitarsis, holotype (NMW), aedeagus, lateral aspect. D. Same, ventral aspect. Scale bars: 0.1 mm.
Fig. 51. Elmomorphus yunnanensis Kodada, Selnekovič & Jäch sp. nov., paratypes (NMW) from type locality (NMW), aedeagus. A. Lateral aspect. B. Lateral aspect. C. Ventral aspect, detail of parameres and penis. Scale bars: 0.1 mm.
Fig. 60. Bursa copulatrix and vagina.A. Elmomorphus nepalensis Satô, 1981, female from Nepal (NMW). B. E. punctulatus sp. nov., paratype from China, locality CWBS 367 (NMW). Scale bar: 0.1 mm.
Fig. 61. Elmomorphus simplipes sp. nov., paratype, male from type locality (CKB), TL: 2.81 mm.
Fig. 63. Elmomorphus donatus Kodada, Selnekovič & Jäch sp. nov., paratype, male from type locality (CKB), TL: 3.30 mm.
Fig. 65. Elmomorphus paradonatus sp. nov., paratype, male from China, locality CWBS 404 (CKB), TL: 3.24 mm.
Fig. 67. Bursa copulatrix and vagina. A. Elmomorphus simplipes sp. nov., paratype from type locality (CKB).B. E. donatus Kodada, Selnekovič & Jäch sp. nov., holotype (NMW), lateral aspect. C. E. donatus, holotype (NMW), ventral aspect. D. E. paradonatus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm.
Fig. 74. Bursa copulatrix and vagina. A. Elmomorphus similis sp. nov., specimen from China, locality CWBS 378 (NMW). B. E. dentipes sp. nov., specimen from China, locality CWBS 354 (NMW). C. E. curvipes sp. nov., specimen from China, locality CWBS 260 (NMW). Scale bar: 0.1 mm.
Fig. 56. Elmomorphus nepalensis Satô, 1981, male from Nepal (NMW), TL: 2.60 mm.
Fig. 57. Elmomorphus nepalensis Satô, 1981, male from Nepal (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 58. Elmomorphus punctulatus sp. nov., paratype, male from China, locality CWBS 367 (CKB), TL: 2.63 mm.
Fig. 59. Elmomorphus punctulatus sp. nov., holotype (IAECAS), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 62. Elmomorphus simplipes sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 64. Elmomorphus donatus Kodada, Selnekovič & Jäch sp. nov., paratype from type locality (CKB), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 66. Elmomorphus paradonatus sp. nov., paratype from China, locality CWBS 404 (CKB), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 69. Elmomorphus similis sp. nov., specimen from China, locality CWBS 354 (CKB), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 71. Elmomorphus dentipes sp. nov., paratype from type locality (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 73. Elmomorphus curvipes sp. nov., paratype from type locality (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 75. Elmomorphus catenatus sp. nov., holotype, male (NMW), TL: 4.14 mm.
Fig. 76. Elmomorphus catenatus sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 78. Elmomorphus jendeki Kodada, Selnekovič & Jäch sp. nov., paratype from type locality (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect, detail of parameres and penis. Scale bar: 0.1 mm.
Fig. 79. Elmomorphus schillhammeri sp. nov., paratype, male from type locality (CKB), TL: 4.26 mm.
Fig. 80. Elmomorphus schillhammeri sp. nov., paratype from type locality (CKB), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 81. Elmomorphus globosus sp. nov., paratype, male from China, locality CWBS 546 (CKB), TL: 3.64 mm.
Fig. 84. Elmomorphus ovalis Kodada, Selnekovič & Jäch sp. nov., paratype from type locality (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 86. Elmomorphus minutus sp. nov., paratype from China, locality CWBS 395 (CKB), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 88. Elmomorphus jii sp. nov., holotype (IAECAS), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 91. A. Elmomorphus schoenmanni sp. nov., holotype (IAECAS), aedeagus, lateral aspect. B. E. vietnamensis sp. nov., paratype from type locality (CKB), aedeagus, lateral aspect. C. Same, ventral aspect. Scale bar: 0.1 mm.
Fig. 92. Elmomorphus reticulatus sp. nov., holotype, male (IAECAS), TL: 3.20 mm.
Fig. 93. Elmomorphus reticulatus sp. nov., paratype from type locality (CKB), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 94. Elmomorphus superficialis sp. nov., paratype, male from type locality (CKB), TL: 3.36 mm.
Fig. 95. Elmomorphus superficialis sp. nov., paratype from type locality (CKB), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 96. Elmomorphus hamatus sp. nov., holotype, male (IAECAS), TL: 3.75 mm.
Fig. 97. Elmomorphus hamatus sp. nov., holotype (IAECAS), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 98. Elmomorphus oblongus sp. nov., holotype, male (NMW), TL: 3.35 mm.
Fig. 99. Elmomorphus oblongus sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 106. Bursa copulatrix and vagina. A. Elmomorphus hamatus sp. nov., paratype from type locality (NMW). B. E. oblongus sp. nov., paratype from type locality (NMW). C. E. glabriclunis sp. nov., paratype from China, locality CWBS 258 (CKB). D. E. simplex sp. nov., paratype from type locality (CKB). E. E. comosiclunis sp. nov., paratype from China, locality CWBS 468. Scale bar: 0.1 mm.
Fig. 103. Elmomorphus simplex sp. nov., paratype from China, locality CWBS 20 (CKB), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.
Fig. 104. Elmomorphus comosiclunis sp. nov., paratype, male from Fujian Province (NMW), TL: 3.69 mm.
Fig. 105. Elmomorphus comosiclunis sp. nov., paratype from Fujian Province (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm.