Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups
FIGURE 1. Some diagnostic characters of the Lepanus australis species group. A, Male protibia of Lepanus eungella dotted line depicts triangular apical digit (grey arrow) and truncate front edge; B, female protibia of Lepanus eungella dotted line depicts truncate front edge, black arrow points to apical spur; C, male protibia of Lepanus australis dotted line depicts elongate apical digit (grey arrow) and deeply angulate front edge; D, female protibia of Lepanus australis dotted line depicts shortened apical digit (grey arrow) and deeply angulate front edge, black arrow points to apical spur; E, lateral view of Lepanus dukungarri white arrow points to row of punctures on metanepisternum; F, lateral view of Lepanus australis, dotted line indicates anteriorly straight inner border of lateral lobe of metaventrite, note the absence of puntures on metanepisternum and straight lower edge of elytral epipleura; G, lateral view of Lepanus oxleyi white arrow points to row of fine punctures on metanepisternum, grey arrow inducates curved lower edge of elytral epipleura; H, male metafemora of Lepanus dukungarri showing medial expansion; I, Head of Lepanus oxleyi white arrow points to interrupted basal carina; J, lateral view of Lepanus latheticus dotted line indicates anteriorly curved inner border of lateral lobe of metaventrite; K, Lepanus australis at bird droppings (image: Geoff Williams).
FIGURE 5. Map of known records including predicted distribution estimated with BIOCLIM models for A, Lepanus australis; B, Lepanus burnetti. QLD, Queensland; NSW, New South Wales; VIC, Victoria.