@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , ; dc:creator "Kosterin, Oleg E.; Onishko, Vladimir V.; Ilyina, Elena V.; Chepurnov, Grigory Yu.; Blinov, Alexander G." ; dc:title "Coenagrion puella" ; dwc:basisOfRecord , , , , , , , , , , , , , , , , , , , , , , , , , , , ; trt:augmentsTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "190" ; bibo:issue "2" ; bibo:journal "Zootaxa" ; bibo:pubDate "2024-06-18" ; bibo:startPage "151" ; bibo:volume "5471" ; dc:creator "Kosterin, Oleg E.; Onishko, Vladimir V.; Ilyina, Elena V.; Chepurnov, Grigory Yu.; Blinov, Alexander G." ; dc:date "2024" ; dc:title "The genus Coenagrion Kirby, 1890 (Odonata: Coenagrionidae) in the Russian part of the Caucasus" ; fabio:hasPart , , , , , ; a fabio:JournalArticle . dwc:authority "(Linnaeus, 1758)" ; dwc:baseAuthorityName "Linnaeus" ; dwc:baseAuthorityYear "1758" ; dwc:class "Insecta" ; dwc:family "Coenagrionidae" ; dwc:genus "Coenagrion" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Linnaeus, 1758)" ; dwc:species "puella" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Coenagrionidae" ; dwc:genus "Coenagrion" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "puella" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Coenagrionidae" ; dwc:genus "Coenagrion" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Coenagrionidae" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 8. Coenagrion puella (a–d) and C. pulchellum (e–m) in different populations in Dagestan in nature; a, j—copulae; b–c—tandems; d, k–m—males; e–g—females; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (178696752); b—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (196674839); c—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag- Khyan Spring, Sirtych, 28.05.2023 (178695642); d —Gergebil’ District, at Gergebil’ Water Reservoir right bank, 24.05.2023 (167302379); e, h, j—Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (178699404, 178699407, 178699406, respectively); f–g, k–l—the same place, 23.05.2023 (166728450, 166734669, 166726653, 166733553, respectively); m—Shuraozen’, 30.05.2023 (181375745); a, c–l—photo by OEK, b, m—by VVO. Not to scale." ; fabio:hasRepresentation ; a fabio:Figure . dwc:eventDate "2019-07-02" ; dwc:locality "Lunnaya Polyana" ; dwc:verbatimElevation "1705" ; dwc:verbatimLatitude "41.156" ; dwc:verbatimLongitude "41.157" ; trt:gbifOccurrenceId "4897992305" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2019-07-03" ; dwc:locality "Forelevoe" ; dwc:verbatimElevation "1264" ; dwc:verbatimLatitude "41.39" ; dwc:verbatimLongitude "41.394" ; trt:gbifOccurrenceId "4897992385" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2019-07-03" ; dwc:locality "Bol'shoy Zelenchuk River" ; dwc:verbatimElevation "1013" ; dwc:verbatimLatitude "41.536" ; dwc:verbatimLongitude "41.54" ; trt:gbifOccurrenceId "4897992350" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2019-07-06" ; dwc:locality "Zelenchuk River" ; dwc:verbatimElevation "580" ; dwc:verbatimLatitude "41.714" ; dwc:verbatimLongitude "41.724" ; trt:gbifOccurrenceId "4897992302" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2019-07-07" ; dwc:locality "Lake Kara-Kel'" ; dwc:verbatimElevation "1340" ; dwc:verbatimLatitude "41.743" ; dwc:verbatimLongitude "41.744" ; trt:gbifOccurrenceId "4897992340" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2019-07-09" ; dwc:locality "Podkumok River" ; dwc:verbatimElevation "1145" ; dwc:verbatimLatitude "43.9" ; dwc:verbatimLongitude "42.386" ; trt:gbifOccurrenceId "4897992359" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2019-07-09" ; dwc:locality "Uchkeken village" ; dwc:verbatimElevation "905" ; trt:gbifOccurrenceId "4897992345" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2023-05-31" ; dwc:locality "Republic" ; dwc:recordedBy "Sarykum Barkhan" ; dwc:verbatimElevation "65" ; dwc:verbatimLatitude "42.9963" ; dwc:verbatimLongitude "47.2201" ; trt:gbifOccurrenceId "4897992362" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Nizhnee Kazanishche village" ; dwc:recordedBy "A. Tatamov" ; dwc:verbatimElevation "512" ; dwc:verbatimLatitude "47.135" ; dwc:verbatimLongitude "47.14" ; trt:gbifOccurrenceId "4897992303" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2023-05-29" ; dwc:locality "Akpeozen' River valley" ; dwc:recordedBy "Buynaksk S" ; dwc:verbatimElevation "488" ; dwc:verbatimLatitude "47.1157" ; dwc:verbatimLongitude "47.1169" ; trt:gbifOccurrenceId "4897992313" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2022-06-22" ; dwc:locality "Lake Gorendzhikor" ; dwc:verbatimElevation "731" ; dwc:verbatimLatitude "46.6758" ; dwc:verbatimLongitude "46.678" ; trt:gbifOccurrenceId "4897992351" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2023-08-05" ; dwc:locality "Lake Bezdonnoe" ; dwc:verbatimElevation "2500" ; dwc:verbatimLatitude "42.5702" ; dwc:verbatimLongitude "46.0162" ; trt:gbifOccurrenceId "4897992376" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2023-07-08" ; dwc:locality "Lake Mochokh" ; dwc:verbatimElevation "1633" ; dwc:verbatimLatitude "42.428" ; dwc:verbatimLongitude "46.524" ; trt:gbifOccurrenceId "4897992355" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2021-06-19" ; dwc:locality "Inchkheozen' River" ; dwc:verbatimElevation "651" ; dwc:verbatimLatitude "47.6703" ; dwc:verbatimLongitude "47.6726" ; trt:gbifOccurrenceId "4897992368" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2023-05-24" ; dwc:locality "Gergebil' District" ; dwc:recordedBy "Gergebil' Water Reservoir" ; dwc:verbatimElevation "810" ; dwc:verbatimLatitude "47.0307" ; dwc:verbatimLongitude "47.0328" ; trt:gbifOccurrenceId "4897992304" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2023-05-28" ; dwc:locality "Rubas River valley" ; dwc:recordedBy "Dzharag-Khyan Spring" ; dwc:verbatimElevation "104" ; dwc:verbatimLatitude "48.2397" ; dwc:verbatimLongitude "48.2411" ; trt:gbifOccurrenceId "4897992377" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2023-05-25" ; dwc:locality "Derbent District" ; dwc:verbatimElevation "18" ; dwc:verbatimLatitude "41.9044" ; dwc:verbatimLongitude "48.4462" ; trt:gbifOccurrenceId "4897992334" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2021-06-25" ; dwc:locality "Bil' - Bil' Kazmalyar village" ; dwc:verbatimElevation "22" ; dwc:verbatimLatitude "48.547" ; dwc:verbatimLongitude "48.548" ; trt:gbifOccurrenceId "4897992378" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Samur Delta Forest" ; dwc:recordedBy "Samurskiy Zakaznik Stationary" ; dwc:verbatimElevation "19" ; dwc:verbatimLatitude "41.8649" ; dwc:verbatimLongitude "48.5541" ; trt:gbifOccurrenceId "4897992331" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2021-06-26" ; dwc:locality "Samur River" ; dwc:verbatimElevation "1370" ; dwc:verbatimLatitude "47.358" ; dwc:verbatimLongitude "47.362" ; trt:gbifOccurrenceId "4897992372" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2019-04-29" ; dwc:locality "Bol'shye Khutora village" ; dwc:recordedBy "Novorossiysk Municipality" ; dwc:verbatimLatitude "44.4782" ; dwc:verbatimLongitude "37.6023" ; trt:gbifOccurrenceId "4897992352" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2018-06-07" ; dwc:locality "Sukko River" ; dwc:recordedBy "Anapa Municipality" ; dwc:verbatimLatitude "37.437" ; dwc:verbatimLongitude "37.439" ; trt:gbifOccurrenceId "4897992301" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2018-06-07" ; dwc:locality "Dzhanhot River" ; dwc:recordedBy "Anapa Municipality" ; dwc:verbatimLatitude "44.459" ; dwc:verbatimLongitude "38.217" ; trt:gbifOccurrenceId "4897992337" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2019-07-04" ; dwc:locality "Bol'shoy Zelenchuk River" ; dwc:verbatimElevation "1014" ; dwc:verbatimLatitude "43.7942" ; dwc:verbatimLongitude "41.5381" ; trt:gbifOccurrenceId "4897992382" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2019-05-01" ; dwc:locality "Kabardino-Balkarian Republic" ; dwc:recordedBy "Mayskiy Town NE" ; dwc:verbatimLatitude "43.654" ; dwc:verbatimLongitude "44.1" ; trt:gbifOccurrenceId "4897992316" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2023-05-24" ; dwc:locality "River" ; dwc:verbatimElevation "559" ; dwc:verbatimLatitude "42.5819" ; dwc:verbatimLongitude "46.9659" ; trt:gbifOccurrenceId "4897992373" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2023-05-28" ; dwc:locality "Rubas River valley" ; dwc:verbatimElevation "180" ; dwc:verbatimLatitude "48.1974" ; dwc:verbatimLongitude "48.1991" ; trt:gbifOccurrenceId "4897992347" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2023-05-27" ; dwc:locality "Bil' - Bil' Kazmalyar village" ; dwc:verbatimElevation "22" ; dwc:verbatimLatitude "41.8075" ; dwc:verbatimLongitude "48.5458" ; trt:gbifOccurrenceId "4897992384" ; trt:httpUri ; a dwc:MaterialCitation .