@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , ; dc:creator "Génier, François; Moretto, Philippe" ; dc:title "Digitonthophagus biflagellatus Genier, new species" ; dwc:basisOfRecord , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; trt:definesTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "110" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:startPage "1" ; bibo:volume "4248" ; dc:creator "Génier, François; Moretto, Philippe" ; dc:date "2017" ; dc:title "Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae)" ; fabio:hasPart , , , , , ; a fabio:JournalArticle . dwc:authority "Genier" ; dwc:authorityName "Genier" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Digitonthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Genier" ; dwc:species "biflagellatus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Digitonthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "biflagellatus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Digitonthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "FIGURES 11 – 14. Male dorsal habitus. 11. D. eucatta, holotype; 12. D. falciger, holotype; 13. Digitonthophagus biflagellatus, holotype; 14. D. gazella, non type (Central Province, Zambia)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 31 – 38. 31, 33, 35, 37. Male head, dorsal view; 32, 34, 36, 38. Male head, frontal view. 31 – 32. Digitonthophagus eucatta, alloreferent; 33 – 34. D. falciger, holotype; 35 – 36. D. biflagellatus, holotype; 37 – 38. D. gazella, non type (Central Province, Zambia)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 55 – 62. Male pronotum, dorsal view. 55. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 56. D. uks, holotype; 57. D. sahelicus, holotype; 58. D. catta, alloreferent (Tamil Nadu State, India); 59. D. eucatta, holotype; 60. D. falciger, holotype; 61. D. biflagellatus, holotype; 62. D. gazella, non type (Central Province, Zambia)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 71 – 78. Female head, dorsal view. 71. Digitonthophagus bonasus, non type (Tamil Nadu State, India); 72. D. uks, allotype; 73. D. sahelicus, allotype; 74. D. catta, non type (India); 75. D. eucatta, allotype; 76. D. falciger, allotype; 77. D. biflagellatus, allotype; 78. D. gazella, non type (Antisiranana Province, Madagascar)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 147 – 158. Internal sac sclerites (FLP, A + SA). 147, 150, 153, 156. Dorsal view; 148, 151, 154, 157. Ventral view. 149, 152, 155, 158. Right lateral view. 147 – 149. Digitonthophagus eucatta, alloreferent (Tamil Nadu State, India); 150 – 152. D. falciger, holotype; 153 – 155. D. biflagellatus, holotype; 156 – 158. D. gazella, non type (Antisiranana Province, Madagascar)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 119 – 134. Parameres, dorsal view. 119. D. bonasus, alloreferent (Tamil Nadu State, India); 120. Digitonthophagus uks, holotype; 121. D. sahelicus, holotype; 122. D. catta, alloreferent (West Bengal, India); 123. D. eucatta, holotype; 124. D. falciger, holotype; 125. D. biflagellatus, holotype; 126. D. gazella, non type (Antisiranana Province, Madagascar); 127. D. petillus, holotype; 128. D. viridicollis, holotype; 129. D. aksumensis, holotype; 130. D. dilatatus, holotype; 131. D. fimator, holotype; 132. D. lusinganus, lectotype; 133. D. ulcerosus, holotype 134. D. namaquensis, holotype." ; fabio:hasRepresentation ; a fabio:Figure . dwc:collectionCode "OUMNH, WORLD" ; dwc:eventDate "1991-05-18" ; dwc:locality "Ndutu area" ; dwc:recordedBy "L. Ndutu & R. Foster" ; dwc:typeStatus "holotype" ; dwc:verbatimLatitude "-2.9833333" ; dwc:verbatimLongitude "35.016666" ; trt:gbifOccurrenceId "1455385224" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "Material" ; dwc:recordedBy "S.R. Penn" ; trt:gbifOccurrenceId "1455385216" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "BURURI" ; dwc:recordedBy "S.R. Penn" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-3.9" ; dwc:verbatimLongitude "29.516666" ; trt:gbifOccurrenceId "1455385241" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IRSNB" ; dwc:eventDate "1940-09-10" ; dwc:locality "Colline Kobero" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-2.649722" ; dwc:verbatimLongitude "30.413334" ; trt:gbifOccurrenceId "1455385232" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OMOC" ; dwc:eventDate "1968-03" ; dwc:locality "RUTANA" ; dwc:recordedBy "S.R. Penn" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2000" ; dwc:verbatimLatitude "-3.75" ; dwc:verbatimLongitude "30.166666" ; trt:gbifOccurrenceId "1455385243" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "Mpinga" ; dwc:recordedBy "S.R. Penn" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-3.75" ; dwc:verbatimLongitude "30.166666" ; trt:gbifOccurrenceId "1455385245" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "PHWC" ; dwc:eventDate "1979-03-26" ; dwc:locality "Parc National Rwindi" ; dwc:recordedBy "SE Parc National Virunga" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-1.3833333" ; dwc:verbatimLongitude "29.466667" ; trt:gbifOccurrenceId "1455385222" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "Rutshuru" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-1.1833334" ; dwc:verbatimLongitude "29.45" ; trt:gbifOccurrenceId "1455385247" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "1952-01-30" ; dwc:locality "Bukavu" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-2.5" ; dwc:verbatimLongitude "28.866667" ; trt:gbifOccurrenceId "1455385242" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "HOMA BAY" ; dwc:recordedBy "Baie de Kavirondo" ; trt:gbifOccurrenceId "1455385233" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:eventDate "1903-09" ; dwc:locality "Lac Victoria" ; dwc:recordedBy "Baie de Kavirondo" ; dwc:verbatimLatitude "-0.51666665" ; dwc:verbatimLongitude "34.483334" ; trt:gbifOccurrenceId "1455385236" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:eventDate "1911-12" ; dwc:locality "Lac Victoria" ; dwc:recordedBy "Alluaud" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-0.51666665" ; dwc:verbatimLongitude "34.483334" ; trt:gbifOccurrenceId "1455385212" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IRSNB" ; dwc:eventDate "1903-10" ; dwc:locality "Rusinga" ; dwc:recordedBy "Victoria-Nyanza N." ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-0.4" ; dwc:verbatimLongitude "34.183334" ; trt:gbifOccurrenceId "1455385274" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BDGC" ; dwc:eventDate "1990-12-23" ; dwc:locality "Nairobi" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1200" ; dwc:verbatimLatitude "-1.75" ; dwc:verbatimLongitude "36.35" ; trt:gbifOccurrenceId "1455385262" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "1911-06-18" ; dwc:locality "Ilala" ; dwc:recordedBy "E Mumias" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1372" ; dwc:verbatimLatitude "0.28333333" ; dwc:verbatimLongitude "34.683334" ; trt:gbifOccurrenceId "1455385284" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:locality "A" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455385239" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "KISUMU" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455385251" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:locality "Kisumu" ; dwc:recordedBy "P.S. Pallas" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-0.1" ; dwc:verbatimLongitude "34.766666" ; trt:gbifOccurrenceId "1455385275" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:eventDate "1912-01" ; dwc:locality "Fort Hall" ; dwc:recordedBy "Alluaud" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-0.71666664" ; dwc:verbatimLongitude "37.15" ; trt:gbifOccurrenceId "1455385253" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "1957-04-24" ; dwc:locality "Nairobi" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-1.2833333" ; dwc:verbatimLongitude "36.816666" ; trt:gbifOccurrenceId "1455385256" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "1993-04" ; dwc:locality "Nairobi National Park" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-1.3666667" ; dwc:verbatimLongitude "36.85" ; trt:gbifOccurrenceId "1455385277" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "1903-06" ; dwc:locality "Kamililo" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455385254" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FGIC, MNHN" ; dwc:eventDate "1913-02" ; dwc:locality "Lemek Valey" ; dwc:recordedBy "G.F. Bornemissza" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-1.1" ; dwc:verbatimLongitude "35.383335" ; trt:gbifOccurrenceId "1455385260" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IRSNB, MNHN" ; dwc:eventDate "1904-01" ; dwc:locality "Bura" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-3.0333333" ; dwc:verbatimLongitude "38.316666" ; trt:gbifOccurrenceId "1455385272" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IRSNB, MNHN" ; dwc:eventDate "1904-01" ; dwc:locality "Mwatate" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-3.5" ; dwc:verbatimLongitude "38.366665" ; trt:gbifOccurrenceId "1455385255" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BDGC" ; dwc:eventDate "1990-12-26" ; dwc:locality "Mwatate" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-3.5" ; dwc:verbatimLongitude "38.366665" ; trt:gbifOccurrenceId "1455385248" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FGIC, MNHN" ; dwc:locality "Mont-Elgon" ; dwc:recordedBy "Foret de Kakamegoes & Environs de Kaimosi" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2000" ; dwc:verbatimLatitude "0.16666667" ; dwc:verbatimLongitude "34.866665" ; trt:gbifOccurrenceId "1455385240" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "PSCW" ; dwc:eventDate "1987-03" ; dwc:locality "Ranch Mpanga" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-2.053611" ; dwc:verbatimLongitude "30.783333" ; trt:gbifOccurrenceId "1455385268" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IRSNB" ; dwc:eventDate "1941-08-21" ; dwc:locality "Bugarula District" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-1.55" ; dwc:verbatimLongitude "29.7" ; trt:gbifOccurrenceId "1455385258" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "Rwaza" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-1.7" ; dwc:verbatimLongitude "29.266666" ; trt:gbifOccurrenceId "1455385271" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FGIC, IRSNB" ; dwc:eventDate "1945-04" ; dwc:locality "Rukwa" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455385269" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IRSNB" ; dwc:eventDate "1948-12" ; dwc:locality "Rukwa" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455385244" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "ARUSHA" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455385270" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "PMOC" ; dwc:eventDate "1997-04-03" ; dwc:locality "Arusha" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-3.3666668" ; dwc:verbatimLongitude "36.683334" ; trt:gbifOccurrenceId "1455385266" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BDGC" ; dwc:eventDate "1990-11-15" ; dwc:locality "Serengeti" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-3.0166667" ; dwc:verbatimLongitude "35.0" ; trt:gbifOccurrenceId "1455385257" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMF" ; dwc:eventDate "1938-03-08" ; dwc:locality "Mumba Hills" ; dwc:recordedBy "Dr. S. G." ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-3.5" ; dwc:verbatimLongitude "35.05" ; trt:gbifOccurrenceId "1455385261" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMF" ; dwc:eventDate "1938-03-23" ; dwc:locality "Mumba Hills" ; dwc:recordedBy "Dr. S. G." ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-3.5" ; dwc:verbatimLongitude "35.05" ; trt:gbifOccurrenceId "1455385237" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMF" ; dwc:eventDate "1938-03-24" ; dwc:locality "Mumba Hills" ; dwc:recordedBy "Dr. S. G." ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-3.5" ; dwc:verbatimLongitude "35.05" ; trt:gbifOccurrenceId "1455385298" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "2012-06-08" ; dwc:locality "Mount Longido" ; dwc:recordedBy "Smith & Takano" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-2.7297223" ; dwc:verbatimLongitude "36.72389" ; trt:gbifOccurrenceId "1455385286" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMNS" ; dwc:eventDate "2007-12-15" ; dwc:locality "DODOMA" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1100" ; dwc:verbatimLatitude "-5.9" ; dwc:verbatimLongitude "35.75" ; trt:gbifOccurrenceId "1455385308" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMNS" ; dwc:eventDate "2006-12-17" ; dwc:locality "Dodoma" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1330" ; dwc:verbatimLatitude "-5.6666665" ; dwc:verbatimLongitude "35.8" ; trt:gbifOccurrenceId "1455385263" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "1917-10" ; dwc:locality "Lulanguru" ; dwc:recordedBy "G. Dellacasa" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1148" ; dwc:verbatimLatitude "-5.1" ; dwc:verbatimLongitude "32.633335" ; trt:gbifOccurrenceId "1455385276" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:locality "Tosamaganga" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-6.6666665" ; dwc:verbatimLongitude "36.316666" ; trt:gbifOccurrenceId "1455385299" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MHNL" ; dwc:eventDate "2011-01-07" ; dwc:locality "Ulembwe Village" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2048" ; dwc:verbatimLatitude "-9.306483" ; dwc:verbatimLongitude "34.620216" ; trt:gbifOccurrenceId "1455385278" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CNC" ; dwc:eventDate "1962-05" ; dwc:locality "Same" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-4.0666666" ; dwc:verbatimLongitude "37.75" ; trt:gbifOccurrenceId "1455385280" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:eventDate "1904-01" ; dwc:locality "Zone" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455385307" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMNS" ; dwc:eventDate "2006-12-20" ; dwc:locality "MANYARA" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1650" ; dwc:verbatimLatitude "-4.5166664" ; dwc:verbatimLongitude "35.35" ; trt:gbifOccurrenceId "1455385279" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "2011-11-22" ; dwc:locality "Mount Hanang" ; dwc:recordedBy "R. Smith" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1964" ; dwc:verbatimLatitude "-4.403889" ; dwc:verbatimLongitude "35.415558" ; trt:gbifOccurrenceId "1455385285" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SMF" ; dwc:locality "Shirati" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-1.15" ; dwc:verbatimLongitude "34.033333" ; trt:gbifOccurrenceId "1455385300" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FETC" ; dwc:eventDate "1966-10-21" ; dwc:locality "Llonga" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-9.033334" ; dwc:verbatimLongitude "36.85" ; trt:gbifOccurrenceId "1455385281" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "ZMUC" ; dwc:eventDate "2004-03-14" ; dwc:locality "Majawanga" ; dwc:recordedBy "S. Tind" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1400" ; dwc:verbatimLatitude "-6.0833335" ; dwc:verbatimLongitude "36.833332" ; trt:gbifOccurrenceId "1455385267" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FGIC, IRSNB" ; dwc:eventDate "1950-12-15" ; dwc:locality "Tumba" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-7.116667" ; dwc:verbatimLongitude "31.166666" ; trt:gbifOccurrenceId "1455385265" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "SHINYANGA" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455385294" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OUMNH" ; dwc:eventDate "1990-11-15" ; dwc:locality "Serengeti National Park" ; dwc:typeStatus "allotype" ; dwc:verbatimLatitude "-2.9833333" ; dwc:verbatimLongitude "35.016666" ; trt:gbifOccurrenceId "1455385282" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OUMNH" ; dwc:eventDate "1990-12-17" ; dwc:locality "Serengeti National Park" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-2.9833333" ; dwc:verbatimLongitude "35.016666" ; trt:gbifOccurrenceId "1455385296" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OUMNH" ; dwc:eventDate "1990-12-18" ; dwc:locality "Serengeti National Park" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-2.9833333" ; dwc:verbatimLongitude "35.016666" ; trt:gbifOccurrenceId "1455385292" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OUMNH" ; dwc:eventDate "1990-12-22" ; dwc:locality "Serengeti National Park" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-2.9833333" ; dwc:verbatimLongitude "35.016666" ; trt:gbifOccurrenceId "1455385273" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OUMNH" ; dwc:eventDate "1990-12-26" ; dwc:locality "Serengeti National Park" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-2.9833333" ; dwc:verbatimLongitude "35.016666" ; trt:gbifOccurrenceId "1455385295" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OUMNH" ; dwc:eventDate "1991-05-18" ; dwc:locality "Serengeti National Park" ; dwc:typeStatus "holotype" ; dwc:verbatimLatitude "-2.9833333" ; dwc:verbatimLongitude "35.016666" ; trt:gbifOccurrenceId "1455385290" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "1935-12-01" ; dwc:locality "Shinyanga" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-3.5666666" ; dwc:verbatimLongitude "33.416668" ; trt:gbifOccurrenceId "1455385283" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "1936-01-26" ; dwc:locality "Shinyanga" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-3.5666666" ; dwc:verbatimLongitude "33.416668" ; trt:gbifOccurrenceId "1455385288" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "BMNH" ; dwc:eventDate "1936-04-10" ; dwc:locality "Shinyanga" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "-3.5666666" ; dwc:verbatimLongitude "33.416668" ; trt:gbifOccurrenceId "1455385264" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; 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