@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , , ; dc:creator "Grisales-Martínez, Freddy A.; Velasco, Julián A.; Bolívar, Wilmar; Williams, Ernest E.; Daza, Juan M." ; dc:title "Anolis urraoi Grisales-Martínez & Velasco & Bolívar & Williams & Daza 2017, sp. nov." ; dwc:basisOfRecord , , , , , , ; trt:citesTaxonName , ; trt:definesTaxonConcept ; trt:deprecates , , , , , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "230" ; bibo:issue "2" ; bibo:journal "Zootaxa" ; bibo:startPage "213" ; bibo:volume "4303" ; dc:creator "Grisales-Martínez, Freddy A.; Velasco, Julián A.; Bolívar, Wilmar; Williams, Ernest E.; Daza, Juan M." ; dc:date "2017" ; dc:title "The taxonomic and phylogenetic status of some poorly known Anolis species from the Andes of Colombia with the description of a nomen nudum taxon" ; fabio:hasPart , , , , , , ; a fabio:JournalArticle . dwc:authority "Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" ; dwc:authorityName "Grisales-Martínez & Velasco & Bolívar & Williams & Daza" ; dwc:authorityYear "2017" ; dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" ; dwc:species "urraoi" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "D'Orbigny" ; dwc:authorityYear "1837" ; dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "D'Orbigny, 1837" ; dwc:species "fuscoauratus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Werner" ; dwc:authorityYear "1916" ; dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Werner, 1916" ; dwc:species "tolimensis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Boulenger" ; dwc:authorityYear "1898" ; dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Boulenger, 1898" ; dwc:species "maculiventris" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Ayala & Williams" ; dwc:authorityYear "1988" ; dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ayala & Williams, 1988" ; dwc:species "medemi" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Cope 1875" ; dwc:authorityName "Cope" ; dwc:authorityYear "1875" ; dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Cope, 1875" ; dwc:species "trachyderma" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Cope 1875" ; dwc:authorityName "Cope" ; dwc:authorityYear "1875" ; dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Cope, 1875" ; dwc:species "bocourti" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Boulenger 1908" ; dwc:authorityName "Boulenger" ; dwc:authorityYear "1908" ; dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Boulenger, 1908" ; dwc:species "scapularis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "urraoi" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Chordata" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "fuscoauratus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "tolimensis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "antonii" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "mariarum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "maculiventris" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "medemi" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "trachyderma" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "bocourti" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:family "Dactyloidae" ; dwc:genus "Anolis" ; dwc:kingdom "Animalia" ; dwc:order "Squamata" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "scapularis" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIGURE 1. Anolis urraoi sp. nov. male holotype (MHUA-R 12733). Photograph by F. Grisales-Martínez." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 3. Holotype of Anolis urraoi sp. nov. (MHUA-R 12733, adult male). A, dorsal; B, lateral; C, ventral view of head; D, detail of dorsal scales at mid-dorsum of body; E, detail of flank scales at mid-dorsum of body; and F, detail of ventral scales at mid-dorsum of body. Scale bar = 5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 4. Dewlap coloration in Andean anoles. A, Anolis urraoi sp. nov., paratype, MHUA-R 12732, photo by F. A. Grisales-Martínez; B, A. urraoi sp. nov., holotype, MHUA-R 12733, photo by F. A. Grisales-Martínez; C, A. urraoi sp. nov., paratype, MHUA-R 12541, photo by F. A. Duarte; D, A. fuscoauratus, QCAZ 8326, Pastaza, Ecuador, photo by M. R. Bustamante, FaunaWebEcuador; E, A. antonii, El Dovio, Valle del Cauca, photo by D. Alzate-Estrada; F, A. antonii, Yotoco, Valle del Cauca, photo by W. Bolívar; G, A. tolimensis, MHUA-R 12770, photo by F. A. Grisales-Martínez; H, A. mariarum MHUA-R 13033, photo by F. A. Grisales-Martínez; and I, A. mariarum MHUA-R 12730, photo by F. A. Grisales-Martínez." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 5. Distribution map of the species in the Anolis fuscoauratus group included in this study. S and M indicate sequenced specimens and measured specimens respectively." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 6. Habitat of Anolis urraoi sp. nov. on the banks of the Penderisco river, Urrao, Antioquia, Colombia." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 7. Maximum likelihood tree (- lnL = - 13314.977) obtained with IQ-TREE depicting the phylogenetic relationships in the Andean species of the Anolis fuscoauratus species group. Inference based on 1465 characters of the ND 2 gene and intervening regions. Nodal support was estimated using the ultrafast bootstrap method using 10000 pseudoreplicates (Minh et al., 2013)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 8. PCA plot of morphometric measurements from specimens of Anolis urraoi sp. nov. (red squares), A. mariarum (magenta circles), A. antonii (blue triangles), A. tolimensis (green diamonds) and some individuals tentatively assigned to A. mariarum (from Western Cordillera at Frontino National Park; black circles). PCA was performed with SVL (snout-vent length) and 22 morphological traits corrected for size (main text for details)." ; fabio:hasRepresentation ; a fabio:Figure . dwc:catalogNumber "MHUA-R 12733" ; dwc:collectionCode "MHUA-R" ; dwc:eventDate "1931-08" ; dwc:locality "Rio Penderisco" ; dwc:recordedBy "B. Rendon-Valencia & Grisales-Martinez & Figs & Table" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "1822" ; trt:gbifOccurrenceId "1585968388" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "MHUA-R 12519-20-21" ; dwc:collectionCode "MHUA-R" ; dwc:locality "Urrao" ; dwc:recordedBy "El Maravillo & Daza" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2120" ; trt:gbifOccurrenceId "1585968389" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "MHUA-R 12521, 12523" ; dwc:collectionCode "MHUA-R" ; dwc:locality "Urrao" ; dwc:recordedBy "El Maravillo & Daza" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "2156" ; trt:gbifOccurrenceId "1585968404" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MHUA-R" ; dwc:eventDate "1931-08" ; dwc:locality "river Penderisco" ; dwc:recordedBy "Urrao & Las Orquideas National Park & Calles & M. A. Serna & Echeverri" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "1816" ; trt:gbifOccurrenceId "1585968385" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Magdalena" ; dwc:recordedBy "Urrao & M. A. Serna" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1585968398" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Geographic" ; dwc:verbatimElevation "2220" ; trt:gbifOccurrenceId "1585968392" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "MHUA-R 12731-32-33" ; dwc:collectionCode "MHUA-R" ; dwc:eventDate "2014-10-31" ; dwc:locality "Specimens" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "1585968416" ; trt:httpUri ; a dwc:MaterialCitation .