@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .
@prefix xlink: <http://www.w3.org/1999/xlink/> .

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    dc:creator "Shapovalov, Andrey M.; Bagaturov, Mikhail F." ;
    dc:title "Lethrus (Mesolethrus) microbuccis Ballion 1871" ;
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    trt:augmentsTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Lethrus_microbuccis_Ballion_1871> ;
    trt:citesTaxonName <http://taxon-name.plazi.org/id/Animalia/Lethrus_bituberculatus> ;
    trt:deprecates <http://taxon-concept.plazi.org/id/Animalia/Lethrus_puncticollis_Kraatz_1882>, <http://taxon-concept.plazi.org/id/Animalia/Lethrus_microbuccis_Semenov_1892>, <http://taxon-concept.plazi.org/id/Animalia/Lethrus_glasunowi_Koshantschikov_1894>, <http://taxon-concept.plazi.org/id/Animalia/Lethrus_microbuccis_Semenov_1894>, <http://taxon-concept.plazi.org/id/Animalia/Lethrus_microbuccis_Nikolajev_1987>, <http://taxon-concept.plazi.org/id/Animalia/Lethrus_microbuccis_Nikolajev_2003> ;
    trt:publishedIn <http://dx.doi.org/10.11646/zootaxa.5351.4.2> ;
    a trt:Treatment .

<http://dx.doi.org/10.11646/zootaxa.5351.4.2>
    bibo:endPage "452" ;
    bibo:issue "4" ;
    bibo:journal "Zootaxa" ;
    bibo:pubDate "2023-09-28" ;
    bibo:startPage "435" ;
    bibo:volume "5351" ;
    dc:creator "Shapovalov, Andrey M.; Bagaturov, Mikhail F." ;
    dc:date "2023" ;
    dc:title "A review of the subgenus Mesolethrus Nikolajev, 2003 of the genus Lethrus Scopoli, 1777 (Coleoptera: Geotrupidae: Lethrinae) with description of a new species" ;
    fabio:hasPart <http://dx.doi.org/10.5281/zenodo.8392021>, <http://dx.doi.org/10.5281/zenodo.8392025>, <http://dx.doi.org/10.5281/zenodo.8392027>, <http://dx.doi.org/10.5281/zenodo.8392033>, <http://dx.doi.org/10.5281/zenodo.8392019>, <http://dx.doi.org/10.5281/zenodo.8392031> ;
    a fabio:JournalArticle .

<http://taxon-concept.plazi.org/id/Animalia/Lethrus_microbuccis_Ballion_1871>
    dwc:authority "Ballion, 1871", "Ballion, 1871: 336" ;
    dwc:authorityName "Ballion" ;
    dwc:authorityPageNumber "336" ;
    dwc:authorityYear "1871" ;
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Ballion, 1871" ;
    dwc:species "microbuccis" ;
    dwc:subGenus "Mesolethrus" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/724FF> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Lethrus_microbuccis> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Lethrus_puncticollis_Kraatz_1882>
    dwc:authority "Kraatz, 1882: 312" ;
    dwc:authorityName "Kraatz" ;
    dwc:authorityPageNumber "312" ;
    dwc:authorityYear "1882" ;
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Kraatz, 1882" ;
    dwc:species "puncticollis" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/724FF> ;
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<http://taxon-concept.plazi.org/id/Animalia/Lethrus_microbuccis_Semenov_1892>
    dwc:authority ": Semenov 1892: 237" ;
    dwc:authorityName ": Semenov" ;
    dwc:authorityPageNumber "237" ;
    dwc:authorityYear "1892" ;
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Semenov, 1892" ;
    dwc:species "microbuccis" ;
    dwc:subGenus "Autolethrus" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/724FF> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Lethrus_microbuccis> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Lethrus_glasunowi_Koshantschikov_1894>
    dwc:authority "D. Koshantschikov, 1894: 118" ;
    dwc:authorityName "D. Koshantschikov" ;
    dwc:authorityPageNumber "118" ;
    dwc:authorityYear "1894" ;
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Koshantschikov, 1894" ;
    dwc:species "glasunowi" ;
    dwc:subGenus "Abrognathus" ;
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    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Lethrus_glasunowi> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Lethrus_microbuccis_Semenov_1894>
    dwc:authority ": Semenov 1894: 479" ;
    dwc:authorityName "Semenov" ;
    dwc:authorityPageNumber "479" ;
    dwc:authorityYear "1894" ;
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Semenov, 1894" ;
    dwc:species "microbuccis" ;
    dwc:subGenus "Microlethrus" ;
    rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/724FF> ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Lethrus_microbuccis> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Lethrus_microbuccis_Nikolajev_1987>
    dwc:authority ": Nikolajev 1987: 41" ;
    dwc:authorityName "Nikolajev" ;
    dwc:authorityPageNumber "41" ;
    dwc:authorityYear "1987" ;
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Nikolajev, 1987" ;
    dwc:species "microbuccis" ;
    dwc:subGenus "Furcilethrus" ;
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    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Lethrus_microbuccis> ;
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<http://taxon-concept.plazi.org/id/Animalia/Lethrus_microbuccis_Nikolajev_2003>
    dwc:authority ": Nikolajev 2003: 126" ;
    dwc:authorityName ": Nikolajev" ;
    dwc:authorityPageNumber "126" ;
    dwc:authorityYear "2003" ;
    dwc:class "Insecta" ;
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    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Nikolajev, 2003" ;
    dwc:species "microbuccis" ;
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    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Lethrus_microbuccis> ;
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<http://taxon-name.plazi.org/id/Animalia/Lethrus_microbuccis>
    dwc:class "Insecta" ;
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    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
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    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Lethrus_Mesolethrus>
    dwc:class "Insecta" ;
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    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "subGenus" ;
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    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Lethrus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Lethrus>
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
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    dwc:order "Coleoptera" ;
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    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Geotrupidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Geotrupidae>
    dwc:class "Insecta" ;
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    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "family" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Coleoptera> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Coleoptera>
    dwc:class "Insecta" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "order" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Insecta> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Insecta>
    dwc:class "Insecta" ;
    dwc:kingdom "Animalia" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "class" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Arthropoda> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Arthropoda>
    dwc:kingdom "Animalia" ;
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    dwc:rank "phylum" ;
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    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia>
    dwc:kingdom "Animalia" ;
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<http://taxon-name.plazi.org/id/Animalia/Lethrus_puncticollis>
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
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    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Lethrus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Lethrus_Autolethrus>
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "subGenus" ;
    dwc:subGenus "Autolethrus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Lethrus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Lethrus_glasunowi>
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:species "glasunowi" ;
    dwc:subGenus "Abrognathus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Lethrus_Abrognathus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Lethrus_Abrognathus>
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "subGenus" ;
    dwc:subGenus "Abrognathus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Lethrus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Lethrus_Microlethrus>
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "subGenus" ;
    dwc:subGenus "Microlethrus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Lethrus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Lethrus_Furcilethrus>
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "subGenus" ;
    dwc:subGenus "Furcilethrus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Lethrus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Lethrus_bituberculatus>
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:species "bituberculatus" ;
    dwc:subGenus "Paralethrus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Lethrus_Paralethrus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Lethrus_Paralethrus>
    dwc:class "Insecta" ;
    dwc:family "Geotrupidae" ;
    dwc:genus "Lethrus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "subGenus" ;
    dwc:subGenus "Paralethrus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Lethrus> ;
    a dwcFP:TaxonName .

<http://dx.doi.org/10.5281/zenodo.8392021>
    dc:description "FIGURES 5–8. Lethrus (Mesolethrus) species, habitus. 5, L. microbuccis Ballion, 1871 (9 km S Ai-Kol); 6, L. elisae Nikolajev, 2001 (Shurob); 7–8, L. dyonini A. Shapovalov & Bagaturov, new species (environs of Tashsay), 7—holotype, 8—allotype. 5–7—males, 8—female." ;
    fabio:hasRepresentation <https://zenodo.org/record/8392021/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.8392025>
    dc:description "FIGURES 9–31. Lethrus (Mesolethrus) species, males genitalia. 9–10, 22–23, L. inermis Reitter, 1897 (9–10—20 km W Oqmachit, 22–23—Pashtamazar); 11–12, 24, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 13–14, 25–27, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 15–16, 28, L. microbuccis Ballion, 1871 (Golbo env.); 17–18, 29, L. elisae Nikolajev, 2001 (Shurob); 19–21, 30–31, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 9–21 aedeagus (9, 11, 13, 15, 17, 19–20—ventral view; 10, 12, 14, 16, 18, 21—lateral view), 22–31 sclerotized apex of endophallus (22, 24, 25, 28–30 dorsal view, 23, 26, 31—lateral view, 27—ventral view)." ;
    fabio:hasRepresentation <https://zenodo.org/record/8392025/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.8392027>
    dc:description "FIGURES 32–43. Lethrus (Mesolethrus) species. 32, 39, L. inermis Reitter, 1897 (Dahanakiik); 33, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 34, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin); 35, 40–42, L. microbuccis Ballion, 1871 (35, 41—Golbo env., 39—Mullabulak, 40—20 km S Guldara); 36, L. elisae Nikolajev, 2001 (Shurob); 37–38, 40–41, L. dyonini A. Shapovalov & Bagaturov, new species, paratypes (environs of Tashsay). 32–37 gena, 38 left mandible, 39 abdomen (lateral view), 40–41 abdomen and apices of elytra (antero-ventral view; 40—male, 41—female), 42–43 tuft of long erect setae on subapical ventrite (lateral view; 42—male, 43—female)." ;
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<http://dx.doi.org/10.5281/zenodo.8392033>
    dc:description "FIGURE 45. Lethrus (Mesolethrus) microbuccis Ballion, 1871, habitus (Kyrgyzstan, between Isfana and Golbo; photograph in nature by D. Milko)." ;
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<http://dx.doi.org/10.5281/zenodo.8392019>
    dc:description "FIGURES 1–4. Lethrus (Mesolethrus) species, male, habitus. 1–2, L. inermis Reitter, 1897 (1—Dahanakiik; 2—20 km W Oqmachit); 3, L. shakhristanicus Nikolajev, 2003 (9 km S Ai-Kol); 4, L. nuratavicus Nikolajev, 1987 (28 km SW Zaamin)." ;
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<http://dx.doi.org/10.5281/zenodo.8392031>
    dc:description "FIGURE 44. Map of distribution Lethrus (Mesolethrus) species: 1, Tashsay env.; 2, south from Belovutsoy River; 3, Khujand; 4, Golbo; 5, 9 km S Ai-Kol; 6, Shahriston env.; 7, Zaamin env.; 8, 28 km SW Zaamin; 9, Sudzhina; 10, Mullabulak; 11, Asmansay; 12, Jangiqishlak env.; 13, 11 km southeast of Djush; 14, Samarkand; 15, 65 km on Samarkand to Qarshi road; 16, Qaynarbuloq; 17, Ruskishlak; 18, 30 km East to Kitab, Qashqadaryo Riv.; 19, Qamar; 20, Guldara; 21, Qarshi; 22, Gumbulok; 23, Okravot Pass; 24, Shurob; 25, Boysun; 26, 20 km W Oqmachit; 27, Chagam; 28, Oqmachit; 29, Shahrinav; 30, Dushanbe env.; 31, Dahanakiik; 32, Yavon; 33, Chormagzak Pass.; 34, Fayzobod; 35, Nurek; 36, Pashtamazar; 37, Kulob. 1–2, 7–8, 10–28 Uzbekistan; 4–5 Kyrgyzstan, 3, 6, 9, 29–37 Tajikistan. Red circles—localities of L. dyonini A. Shapovalov & Bagaturov, new species (1—type locality: Tashsay); circles with green colour—localities of L. microbuccis Ballion, 1871 (3—type locality: Khujand); circles with white colour—localities of L. shakhristanicus Nikolajev, 2003 (6—type locality: Shahriston env.); circles with blue colour—localities of L. nuratavicus Nikolajev, 1987 (11—type locality: Asmansay); circles with yellow colour— localities of L. elisae Nikolajev, 2001 (24—type locality: Shurob); circles with orange colour—localities of L. inermis Reitter, 1897 (37—type locality: Kulab?); orange asterisk—Kyrgyzstan, Taldyk (erroneous record of L. inermis from Protzenko 1968)." ;
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