@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , ; dc:creator "Grischenko, Andrei V.; Gordon, Dennis P.; Morozov, Taras B." ; dc:title "Fatkullina imitata Grischenko & Gordon & Morozov 2018, n. sp." ; dwc:basisOfRecord , , , , , , , , , , ; trt:definesTaxonConcept ; trt:publishedIn ; a trt:Treatment . dc:creator "Grischenko, Andrei V.; Gordon, Dennis P.; Morozov, Taras B." ; dc:date "2018" ; dc:title "Fatkullina imitata n. sp., second species of a unique cheilostome bryozoan genus with reversed-polarity zooidal budding, and new family Fatkullinidae" ; fabio:hasPart , , , , , . dwc:authority "Grischenko & Gordon & Morozov, 2018" ; dwc:authorityName "Grischenko & Gordon & Morozov" ; dwc:authorityYear "2018" ; dwc:class "Gymnolaemata" ; dwc:family "Stomachetosellidae" ; dwc:genus "Fatkullina" ; dwc:kingdom "Animalia" ; dwc:order "Cheilostomatida" ; dwc:phylum "Bryozoa" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Grischenko & Gordon & Morozov, 2018" ; dwc:species "imitata" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Gymnolaemata" ; dwc:family "Stomachetosellidae" ; dwc:genus "Fatkullina" ; dwc:kingdom "Animalia" ; dwc:order "Cheilostomatida" ; dwc:phylum "Bryozoa" ; dwc:rank "species" ; dwc:species "imitata" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Gymnolaemata" ; dwc:family "Stomachetosellidae" ; dwc:genus "Fatkullina" ; dwc:kingdom "Animalia" ; dwc:order "Cheilostomatida" ; dwc:phylum "Bryozoa" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Gymnolaemata" ; dwc:family "Stomachetosellidae" ; dwc:kingdom "Animalia" ; dwc:order "Cheilostomatida" ; dwc:phylum "Bryozoa" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Gymnolaemata" ; dwc:kingdom "Animalia" ; dwc:order "Cheilostomatida" ; dwc:phylum "Bryozoa" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Gymnolaemata" ; dwc:kingdom "Animalia" ; dwc:phylum "Bryozoa" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Bryozoa" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "FIGURE 1. General view of dried colonies of Fatkullina imitata n. sp. (A–D) and F. paradoxa Grischenko, Gordon & Taylor, 1998 (E). A, B, specimens R.V. Professor Kizevetter 2015, Stn 73; C, D, specimens paratype 8 NIWA 127751; E, specimen R.V. Professor Probatov 2013, Stn 5, 16 June 2013, 57.41694° N, 156.51806° E, 27 m, pebbles, sand, silt. A, B. Colonies encrusting pebbles. C, D. Colonies encrusting external (C), and internal (D) surface of shell fragment of Chlamys sp. E. Colony encrusting internal surface of bivalve shell fragment of Serripes sp., surrounded by other encrusting bryozoan species. Scale bars: 5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 2. Fatkullina imitata n. sp. A, Holotype ZIRAS 1/50661; B, paratype 5 ZIRAS 6/50666; C, paratype 4 ZIRAS 5/ 50665. A. General view of colony encrusting flattened pebble in boundary interactions with Celleporina ventricosa (Lorenz, 1886) (top left), Reussinella arctica (Osburn, 1950) (top right), Hippothoa arctica (Kluge, 1906), H. expansa Dawson, 1859 (bottom left) and Pachyegis princeps (Norman, 1903) (centrally to bottom right). B. General view of a medium-sized colony. C. General view of a young colony demonstrating a gradual shift in geometry of zooidal budding from spiral into reversedpolarity. Asterisks indicate zooids with a sideways (transverse) orientation. Scale bars: 1 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 3. Fatkullina imitata n. sp. A, holotype ZIRAS 1/50661; B–D, paratype 5 ZIRAS 6/50666; E, F, paratype 2 ZIRAS 3/50663. A, B. Zooids differentiating at colony margin. C, D. Randomly orientated autozooids in central colony area. E. Three zooids, showing granular-tubercular frontal walls uniformly perforated by large, round, deep infundibuliform pseudopores and drop-shaped secondary orifices with rugged outlines, overhanging distal curvature of primary orifice and slightly covering its sides. F. Distolateral view towards two zooids, showing roughly granular-tubercular, elevated and distended frontal wall around orificial area, evenly perforated with circular to oval deep infundibuliform pseudopores. Scale bars: A–D, 0.25 mm; E, F, 0.1 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 5. Fatkullina imitata n. sp. A, paratype 5 ZIRAS 6/50666; B, holotype ZIRAS 1/50661; C–G, paratype 2 ZIRAS 3/ 50663. A. Orificial area of newly formed zooid near colony margin, showing transversely oval primary orifice bordered by thinly calcified smooth narrow rim, and surrounded by granular-tubercular surface with circular pseudopores. B. Orificial area of completed zooid, showing drop-shaped secondary orifice with granular tubercular surface, overhanging distal curvature of deeply submerged primary orifice, leaving visible its proximal thickened smooth shelf with shallow sinus; note circular infundibular pseudopores with finely tuberculated surfaces stretched deep inside. C. Interior of zooidal orificial area, showing transversally oval primary orifice with shallow sinus and deep calcified visor with denticulate margin, covers half of primary orifice length. D. Interior of group of zooids from central colony area, indicating their incoherent orientation, showing transversally oval primary orifices, smaller circular scattered openings of pseudopores in frontal shields, and enlarged circular to oval openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom. E. Interior of frontal shield, showing scattered smaller openings of pseudopores centrally, enlarged openings of tubular areolar pore channels encroaching via vertical walls, which connect main coelom with hypostegal coelom, and transversally oval primary orifice with distal overhanging visor of secondary calcification covering more than 2/3 of primary orifice length. F. Internal and external view of zooidal lateral wall with basal pore chambers and external openings of tubular areolar pore channels encroaching lateral wall to open into the hypostegal coelom. G. Longitudinal cross-section through zooid, showing remarkable thickness of frontal shield, with deep, proximally tilted lumen between primary and secondary orifices, and lateral to frontal openings of tubular areolar pore channels connecting main coelom with hypostegal coelom. Scale bars: A–C, E–G, 0.1 mm; D, 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 4. Fatkullina imitata n. sp. A–C, E, F, paratype 1 ZIRAS 2/50662; D, holotype ZIRAS 1/50661. Intracolonial polymorphism. A. Strongly distended zooids in elevated colony center, with proportionally elongated and elevated distal (preoral) frontal wall terminating with conical and bulbous solid umbones (arrowheads). B. Two neighboring zooids, each bearing distal (preoral) conical knob (arrowheads), with lateral boundary occluded by secondary calcification, indicated by enlarged infundibular pseudopores; note minute shortened distal suture line separating preoral areas. C. Two oppositely orientated zooids with strongly swollen frontal shields and orifices shifted to the top of frontal surface. D. Irregularly hexagonal, flattened, uniformly perforated kenozooid with undulating margins, surrounded by six autozooids. E. Colony area comprising mixed kenozooids and autozooids; note irregular shape of flattened kenozooids possessing strongly reduced orificial areas remaining in form of larger frontal pseudopores (left to central kenozooids) or two minor pseudopores (right kenozooid). F. Colony area fully represented by flattened kenozooids with orifices constricted or closed by calcified cylindrical or mammiform projections with tiny circular to slit-like openings at their apices. Scale bars: 0.25 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 6. Fatkullina imitata n. sp. A–G, paratype 3 ZIRAS 4/50664; H, specimen R.V. TINRO 2016, Stn 74. Early astogeny. A–C, ancestrular complex formation; D–F, ancestrular interior; G, H, colonies at early stages of astogeny, external and internal views. A. Ancestrula with first distally-budded, transversally/spirally arranged postancestrular zooid. B. Ancestrula with two postancestrular zooids (right bud partially damaged) spirally arranged and budded in opposite directions; note cormidial nature of ancestrular orifice, with its distal margin comprises lateral vertical wall of distally budded zooid, and right lateral margin – lateral vertical wall of laterally budded zooid. C. Ancestrula with two postancestrular zooids, spirally arranged and budded in same direction. D. Interior of ancestrula and distally arranged, perpendicularly orientated periancestrular zooid. E. Interior of ancestrular zooid, showing intensive distal umbonuloid area around orifice composed of four lobes separated with fine meandering sutures, ring scar, and proximal pseudoporous (lepralioid) part of the frontal shield. F. Close-up of umbonuloid area including circular orifice possessing deep V-shaped sinus, four uneven lobes separated with fine meandering sutures, discrete ring scar forming regular boundary between exterior-wall microstructure of umbonuloid part and extra-umbonuloid calcification. G. Ancestrular complex comprising ancestrula and three spirally arranged periancestrular zooids plus four developing to fully completed zooids. H. Undersides of ancestrular complex. Scale bars: A–D, G, H, 0.2 mm; E, F, 0.1 mm." ; fabio:hasRepresentation ; a fabio:Figure . dwc:catalogNumber "ZIRAS 1" ; dwc:collectionCode "ZIRAS" ; dwc:eventDate "2008-06-16" ; dwc:locality "Sea of Okhotsk" ; dwc:recordedBy "Collection, R. V & S. G. Korostylyov." ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "290" ; dwc:verbatimLatitude "58.03833" ; dwc:verbatimLongitude "155.72028" ; trt:gbifOccurrenceId "2028138719" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ZIRAS 2" ; dwc:collectionCode "ZIRAS" ; dwc:eventDate "2008-06-16" ; dwc:locality "Sea of Okhotsk" ; dwc:recordedBy "Collection, R. V & S. G. Korostylyov." ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "290" ; dwc:verbatimLatitude "58.03833" ; dwc:verbatimLongitude "155.72028" ; trt:gbifOccurrenceId "2028138713" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ZIRAS 3" ; dwc:collectionCode "ZIRAS" ; dwc:eventDate "2015-07-03" ; dwc:locality "Sea of Okhotsk" ; dwc:recordedBy "Collection, R. V. & T. B. Morozov." ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "145" ; dwc:verbatimLatitude "58.13333" ; dwc:verbatimLongitude "156.01889" ; trt:gbifOccurrenceId "2028138720" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ZIRAS 4" ; dwc:collectionCode "ZIRAS" ; dwc:eventDate "2015-07-03" ; dwc:locality "Sea of Okhotsk" ; dwc:recordedBy "Collection, R. V. & T. B. Morozov." ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "145" ; dwc:verbatimLatitude "58.13333" ; dwc:verbatimLongitude "156.01889" ; trt:gbifOccurrenceId "2028138723" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ZIRAS 5" ; dwc:collectionCode "ZIRAS" ; dwc:eventDate "2008-06-16" ; dwc:locality "Sea of Okhotsk" ; dwc:recordedBy "Collection, R. V & S. G. Korostylyov." ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "290" ; dwc:verbatimLatitude "58.03833" ; dwc:verbatimLongitude "155.72028" ; trt:gbifOccurrenceId "2028138716" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ZIRAS 6" ; dwc:collectionCode "ZIRAS" ; dwc:eventDate "2008-06-16" ; dwc:locality "Sea of Okhotsk" ; dwc:recordedBy "Collection, R. V & S. G. Korostylyov." ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "285" ; dwc:verbatimLatitude "58.01917" ; dwc:verbatimLongitude "155.71667" ; trt:gbifOccurrenceId "2028138714" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "NHMUK 2017.7" ; dwc:collectionCode "NHMUK" ; dwc:eventDate "2008-06-16" ; dwc:locality "Sea of Okhotsk" ; dwc:recordedBy "Collection, R. V & S. G. Korostylyov." ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "290" ; dwc:verbatimLatitude "58.03833" ; dwc:verbatimLongitude "155.72028" ; trt:gbifOccurrenceId "2028138722" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "NHMUK 2017.7" ; dwc:collectionCode "NHMUK" ; dwc:eventDate "2013-08-11" ; dwc:locality "Sea of Okhotsk" ; dwc:recordedBy "Collection, R. V. & T. B. Morozov." ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "146" ; dwc:verbatimLatitude "58.15" ; dwc:verbatimLongitude "156.05" ; trt:gbifOccurrenceId "2028138718" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "NIWA 127751" ; dwc:collectionCode "NIWA" ; dwc:eventDate "2013-08-11" ; dwc:locality "Sea of Okhotsk" ; dwc:recordedBy "Collection, R. V & Stn & R. V. Professor Probatov" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "146" ; dwc:verbatimLatitude "58.15" ; dwc:verbatimLongitude "156.05" ; trt:gbifOccurrenceId "2028138721" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "TINRO" ; dwc:recordedBy "Stn & R. V. Professor Kizevetter" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "2028138715" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "TINRO" ; dwc:recordedBy "Stn" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "2028138717" ; trt:httpUri ; a dwc:MaterialCitation .