@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .

<http://treatment.plazi.org/id/03B687B92D7FFFDA76BFFF09FE14A4E1>
    cito:cites <http://dx.doi.org/10.5281/zenodo.5901045>, <http://dx.doi.org/10.5281/zenodo.5901049>, <http://dx.doi.org/10.5281/zenodo.5901053>, <http://dx.doi.org/10.5281/zenodo.5901055>, <http://dx.doi.org/10.5281/zenodo.5901059>, <http://dx.doi.org/10.5281/zenodo.5901063>, <http://dx.doi.org/10.5281/zenodo.5901065>, <http://dx.doi.org/10.5281/zenodo.5901069>, <http://dx.doi.org/10.5281/zenodo.5901071>, <http://taxon-concept.plazi.org/id/Animalia/Bothynus_alvarengai_Martinez_1983>, <http://taxon-concept.plazi.org/id/Animalia/Bothynus_belemensis_Duarte_2022>, <http://taxon-concept.plazi.org/id/Animalia/Bothynus_horridus_Endrodi_1968>, <http://taxon-concept.plazi.org/id/Animalia/Bothynus_spinophallicus_Ratcliffe_2021> ;
    dc:creator "Duarte, Paulo R. M.; Grossi, Paschoal C.; Dupuis, Fabien" ;
    dc:title "Bothynus villiersi Endrodi 1968" ;
    trt:augmentsTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Bothynus_villiersi_Endrodi_1968> ;
    trt:publishedIn <http://dx.doi.org/10.11646/zootaxa.5093.1.3> ;
    a trt:Treatment .

<http://dx.doi.org/10.11646/zootaxa.5093.1.3>
    bibo:endPage "66" ;
    bibo:issue "1" ;
    bibo:journal "Zootaxa" ;
    bibo:pubDate "2022-01-25" ;
    bibo:startPage "49" ;
    bibo:volume "5093" ;
    dc:creator "Duarte, Paulo R. M.; Grossi, Paschoal C.; Dupuis, Fabien" ;
    dc:date "2022" ;
    dc:title "A taxonomic revision of the Bothynus villiersi Endrödi, 1968 species group (Coleoptera: Scarabaeidae: Dynastinae)" ;
    fabio:hasPart <http://dx.doi.org/10.5281/zenodo.5901045>, <http://dx.doi.org/10.5281/zenodo.5901049>, <http://dx.doi.org/10.5281/zenodo.5901053>, <http://dx.doi.org/10.5281/zenodo.5901055>, <http://dx.doi.org/10.5281/zenodo.5901059>, <http://dx.doi.org/10.5281/zenodo.5901063>, <http://dx.doi.org/10.5281/zenodo.5901065>, <http://dx.doi.org/10.5281/zenodo.5901069>, <http://dx.doi.org/10.5281/zenodo.5901071> ;
    a fabio:JournalArticle .

<http://taxon-concept.plazi.org/id/Animalia/Bothynus_villiersi_Endrodi_1968>
    dwc:authority "Endrodi, 1968" ;
    dwc:authorityName "Endrodi" ;
    dwc:authorityYear "1968" ;
    dwc:class "Insecta" ;
    dwc:family "Dynastidae" ;
    dwc:genus "Bothynus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Endrodi, 1968" ;
    dwc:species "villiersi" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Bothynus_villiersi> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Bothynus_horridus_Endrodi_1968>
    dwc:authority "Endrodi, 1968" ;
    dwc:authorityName "Endrodi" ;
    dwc:authorityYear "1968" ;
    dwc:class "Insecta" ;
    dwc:family "Dynastidae" ;
    dwc:genus "Bothynus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Endrodi, 1968" ;
    dwc:species "horridus" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Bothynus_horridus> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Bothynus_alvarengai_Martinez_1983>
    dwc:authority "Martinez, 1983" ;
    dwc:authorityName "Martinez" ;
    dwc:authorityYear "1983" ;
    dwc:class "Insecta" ;
    dwc:family "Dynastidae" ;
    dwc:genus "Bothynus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Martinez, 1983" ;
    dwc:species "alvarengai" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Bothynus_alvarengai> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Bothynus_spinophallicus_Ratcliffe_2021>
    dwc:authority "(Ratcliffe & Cave, 2021)" ;
    dwc:authorityName "Duarte & Grossi & Dupuis" ;
    dwc:authorityYear "2022" ;
    dwc:baseAuthorityName "Ratcliffe & Cave" ;
    dwc:baseAuthorityYear "2021" ;
    dwc:class "Insecta" ;
    dwc:family "Dynastidae" ;
    dwc:genus "Bothynus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Ratcliffe & Cave, 2021)" ;
    dwc:species "spinophallicus" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Bothynus_spinophallicus> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Bothynus_belemensis_Duarte_2022>
    dwc:authority "Duarte, Grossi, & Dupuis" ;
    dwc:authorityName "Duarte, Grossi, & Dupuis" ;
    dwc:authorityYear "2022" ;
    dwc:class "Insecta" ;
    dwc:family "Dynastidae" ;
    dwc:genus "Bothynus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Duarte, Grossi, & Dupuis, 2022" ;
    dwc:species "belemensis" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Bothynus_belemensis> ;
    a dwcFP:TaxonConcept .

<http://taxon-name.plazi.org/id/Animalia/Bothynus_villiersi>
    dwc:class "Insecta" ;
    dwc:family "Dynastidae" ;
    dwc:genus "Bothynus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Coleoptera" ;
    dwc:phylum "Arthropoda" ;
    dwc:rank "species" ;
    dwc:species "villiersi" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Bothynus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Bothynus>
    dwc:genus "Bothynus" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Dynastidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Dynastidae>
    dwc:family "Dynastidae" ;
    dwc:rank "family" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Coleoptera> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Coleoptera>
    dwc:order "Coleoptera" ;
    dwc:rank "order" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Insecta> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Insecta>
    dwc:class "Insecta" ;
    dwc:rank "class" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Arthropoda> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Arthropoda>
    dwc:phylum "Arthropoda" ;
    dwc:rank "phylum" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia>
    dwc:kingdom "Animalia" ;
    dwc:rank "kingdom" ;
    a dwcFP:TaxonName .

<http://dx.doi.org/10.5281/zenodo.5901045>
    dc:description "FIGURE 1. Morphological characters of the Bothynus villiersi species group: A, head in lateral view (mouthparts removed); B, head in frontal view; C, right maxilla in ventral view; D, head, prothoracic legs and pronotum in lateral view. Arrows indicate oblique downward clypeus in A, complete frontoclypeal ridge in B, palpomere 1 short in C, and conical process of mentum in D." ;
    fabio:hasRepresentation <https://zenodo.org/record/5901045/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.5901049>
    dc:description "FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021)." ;
    fabio:hasRepresentation <https://zenodo.org/record/5901049/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.5901053>
    dc:description "FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype)." ;
    fabio:hasRepresentation <https://zenodo.org/record/5901053/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.5901055>
    dc:description "FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D." ;
    fabio:hasRepresentation <https://zenodo.org/record/5901055/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.5901059>
    dc:description "FIGURE 5. Spiculum gastrale: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. hs = hemisternite." ;
    fabio:hasRepresentation <https://zenodo.org/record/5901059/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.5901063>
    dc:description "FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B." ;
    fabio:hasRepresentation <https://zenodo.org/record/5901063/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.5901065>
    dc:description "FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species." ;
    fabio:hasRepresentation <https://zenodo.org/record/5901065/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.5901069>
    dc:description "FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate." ;
    fabio:hasRepresentation <https://zenodo.org/record/5901069/files/figure.png> ;
    a fabio:Figure .

<http://dx.doi.org/10.5281/zenodo.5901071>
    dc:description "FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record." ;
    fabio:hasRepresentation <https://zenodo.org/record/5901071/files/figure.png> ;
    a fabio:Figure .