@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . cito:cites , , , , , , , , , , ; dc:creator "Duarte, Paulo R. M.; Grossi, Paschoal C.; Dupuis, Fabien" ; dc:title "Bothynus alvarengai Martinez 1983" ; dwc:basisOfRecord , , , , , , , , , , , , ; trt:augmentsTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "66" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:pubDate "2022-01-25" ; bibo:startPage "49" ; bibo:volume "5093" ; dc:creator "Duarte, Paulo R. M.; Grossi, Paschoal C.; Dupuis, Fabien" ; dc:date "2022" ; dc:title "A taxonomic revision of the Bothynus villiersi Endrödi, 1968 species group (Coleoptera: Scarabaeidae: Dynastinae)" ; fabio:hasPart , , , , , , , ; a fabio:JournalArticle . dwc:authority "Martinez, 1983" ; dwc:authorityName "Martinez" ; dwc:authorityYear "1983" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Bothynus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Martinez, 1983" ; dwc:species "alvarengai" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Endrodi" ; dwc:authorityYear "1968" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Bothynus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Endrodi, 1968" ; dwc:species "villiersi" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Endrodi" ; dwc:authorityYear "1968" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Bothynus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Endrodi, 1968" ; dwc:species "horridus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Duarte & Grossi & Dupuis, 2022" ; dwc:authorityName "Duarte & Grossi & Dupuis" ; dwc:authorityYear "2022" ; dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Bothynus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Duarte, Grossi & Dupuis, 2022" ; dwc:species "alvarengai" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Dynastidae" ; dwc:genus "Bothynus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "alvarengai" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:genus "Bothynus" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Dynastidae" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:order "Coleoptera" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 5. Spiculum gastrale: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. hs = hemisternite." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record." ; fabio:hasRepresentation ; a fabio:Figure . dwc:collectionCode "MACN" ; dwc:eventDate "1980-12" ; dwc:locality "Encruzilhada" ; dwc:recordedBy "A. Martinez & Alvarenga" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3455072339" ; trt:httpUri ; a dwc:MaterialCitation . dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3455072321" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "EPGC" ; dwc:eventDate "2007-12-11" ; dwc:locality "Encruzilhada" ; dwc:recordedBy "Rafael & Parizotto" ; dwc:verbatimElevation "800" ; dwc:verbatimLatitude "-15.540278" ; dwc:verbatimLongitude "-40.836666" ; trt:gbifOccurrenceId "3455072355" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CZMA" ; dwc:eventDate "2007-11-11" ; dwc:locality "Base de Geraldina" ; dwc:recordedBy "F. Limeira-de-Oliveira" ; trt:gbifOccurrenceId "3455072313" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CERPE" ; dwc:eventDate "2011-04-02" ; dwc:locality "Posto Avancado do Mel" ; dwc:recordedBy "F. Limeira-de-Oliveira & G. A. Reis & M. S. Oliveira" ; dwc:verbatimLatitude "-6.7305555" ; dwc:verbatimLongitude "-44.983055" ; trt:gbifOccurrenceId "3455072308" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CERPE" ; dwc:eventDate "1997-12" ; dwc:locality "Aguas Vermelhas" ; dwc:recordedBy "Vaz-de-Mello & Belo" ; trt:gbifOccurrenceId "3455072352" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CERPE" ; dwc:eventDate "2012-12-12" ; dwc:locality "Fazenda Faceiro" ; dwc:recordedBy "Grossi & P. Grossi. J. A. Rafael & G. A. R. Melo" ; trt:gbifOccurrenceId "3455072356" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "EPGC" ; dwc:eventDate "2007-11-15" ; dwc:locality "Fazenda Veredao" ; dwc:recordedBy "Grossi & Rafael & Parizotto" ; dwc:verbatimLatitude "-15.6647215" ; dwc:verbatimLongitude "-41.665558" ; trt:gbifOccurrenceId "3455072329" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FDPC" ; dwc:eventDate "1993-11" ; dwc:locality "Ipatinga" ; trt:gbifOccurrenceId "3455072369" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "EPGC" ; dwc:eventDate "2011-12-22" ; dwc:locality "Parque Estadual do Rio Preto" ; dwc:recordedBy "Oliveira & Ferreira" ; trt:gbifOccurrenceId "3455072346" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FDPC" ; dwc:eventDate "1992-12" ; dwc:locality "Itatiaia" ; trt:gbifOccurrenceId "3455072343" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Encruzilhada" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "3455072366" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "Maranhao" ; trt:gbifOccurrenceId "3455072340" ; trt:httpUri ; a dwc:MaterialCitation .