@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . cito:cites , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; dc:creator "Kpan, Tokouaho Flora; Kouamé, N’Goran Germain; Barej, Michael F.; Adeba, Patrick Joël; Emmrich, Mike; Ofori-Boateng, Caleb; Rödel, Mark-Oliver" ; dc:title "Phrynobatrachus tanoeensis Kpan & Kouamé & Barej & Adeba & Emmrich & Ofori-Boateng & Rödel 2018, sp. nov." ; dwc:basisOfRecord , , , ; trt:citesTaxonName ; trt:definesTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "237" ; bibo:issue "2" ; bibo:journal "Zootaxa" ; bibo:pubDate "2018-03-02" ; bibo:startPage "221" ; bibo:volume "4388" ; dc:creator "Kpan, Tokouaho Flora; Kouamé, N’Goran Germain; Barej, Michael F.; Adeba, Patrick Joël; Emmrich, Mike; Ofori-Boateng, Caleb; Rödel, Mark-Oliver" ; dc:date "2018" ; dc:title "A new Puddle Frog, genus Phrynobatrachus (Amphibia: Anura: Phrynobatrachidae), from the eastern part of the Upper Guinea biodiversity hotspot, West Africa" ; fabio:hasPart , , , , ; a fabio:JournalArticle . dwc:authority "Kpan & Kouamé & Barej & Adeba & Emmrich & Ofori-Boateng & Rödel, 2018" ; dwc:authorityName "Kpan & Kouamé & Barej & Adeba & Emmrich & Ofori-Boateng & Rödel" ; dwc:authorityYear "2018" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Kpan, Kouamé, Barej, Adeba, Emmrich, Ofori-Boateng & Rödel, 2018" ; dwc:species "tanoeensis" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Gunther" ; dwc:authorityYear "1862" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Gunther, 1862" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Smith" ; dwc:authorityYear "1849" ; dwc:class "Amphibia" ; dwc:family "Arthroleptidae" ; dwc:genus "Arthroleptis" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Smith, 1849" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Boulenger" ; dwc:authorityYear "1917" ; dwc:class "Amphibia" ; dwc:family "Ptychadenidae" ; dwc:genus "Ptychadena" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Boulenger, 1917" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Dubois" ; dwc:authorityYear "1987" ; dwc:class "Amphibia" ; dwc:family "Ptychadenidae" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "family" ; dwc:scientificNameAuthorship "Dubois, 1987" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Barbour & Loveridge" ; dwc:authorityYear "1927" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Barbour & Loveridge, 1927" ; dwc:species "liberiensis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "(Gunther, 1858)" ; dwc:authorityName "Gunther" ; dwc:authorityYear "1858" ; dwc:baseAuthorityName "Gunther" ; dwc:baseAuthorityYear "1858" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Gunther, 1858)" ; dwc:species "plicatus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Perret, 1966" ; dwc:authorityName "Perret" ; dwc:authorityYear "1966" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Perret, 1966" ; dwc:species "annulatus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "(Peters, 1863)" ; dwc:authorityName "Peters" ; dwc:authorityYear "1863" ; dwc:baseAuthorityName "Peters" ; dwc:baseAuthorityYear "1863" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Peters, 1863)" ; dwc:species "calcaratus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Perret, 1988" ; dwc:authorityName "Perret" ; dwc:authorityYear "1988" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Perret, 1988" ; dwc:species "taiensis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Guibe, 1959" ; dwc:authorityName "Guibe" ; dwc:authorityYear "1959" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Guibe, 1959" ; dwc:species "villiersi" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Hillers, Zimkus & Rodel, 2008" ; dwc:authorityName "Hillers, Zimkus & Rodel" ; dwc:authorityYear "2008" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Hillers, Zimkus & Rodel, 2008" ; dwc:species "pintoi" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "(Chabanaud, 1921)" ; dwc:authorityName "Chabanaud" ; dwc:authorityYear "1921" ; dwc:baseAuthorityName "Chabanaud" ; dwc:baseAuthorityYear "1921" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Chabanaud, 1921)" ; dwc:species "gutturosus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Rodel, Ohler & Hillers, 2010" ; dwc:authorityName "Rodel, Ohler & Hillers" ; dwc:authorityYear "2010" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Rodel, Ohler & Hillers, 2010" ; dwc:species "hieroglyphicus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "(Chabanaud, 1921)" ; dwc:authorityName "Chabanaud" ; dwc:authorityYear "1921" ; dwc:baseAuthorityName "Chabanaud" ; dwc:baseAuthorityYear "1921" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Chabanaud, 1921)" ; dwc:species "tokba" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "(Chabanaud, 1921)" ; dwc:authorityName "Chabanaud" ; dwc:authorityYear "1921" ; dwc:baseAuthorityName "Chabanaud" ; dwc:baseAuthorityYear "1921" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Chabanaud, 1921)" ; dwc:species "fraterculus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Schiotz" ; dwc:authorityYear "1964" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Schiotz, 1964" ; dwc:species "ghanensis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Guibe & Lamotte, 1958" ; dwc:authorityName "Guibe & Lamotte" ; dwc:authorityYear "1958" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Guibe & Lamotte, 1958" ; dwc:species "maculiventris" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority ", Ahl, 1924" ; dwc:authorityName ", Ahl" ; dwc:authorityYear "1924" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Ahl, 1924" ; dwc:species "latifrons" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Parker, 1936" ; dwc:authorityName "Parker" ; dwc:authorityYear "1936" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Parker, 1936" ; dwc:species "alleni" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Boulenger, 1912" ; dwc:authorityName "Boulenger" ; dwc:authorityYear "1912" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Boulenger, 1912" ; dwc:species "francisci" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "(Smith, 1849)" ; dwc:authorityName "Smith" ; dwc:authorityYear "1849" ; dwc:baseAuthorityName "Smith" ; dwc:baseAuthorityYear "1849" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Smith, 1849)" ; dwc:species "natalensis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Rodel, Onadeko, Barej & Sandberger, 2012" ; dwc:authorityName "Rodel, Onadeko, Barej & Sandberger" ; dwc:authorityYear "2012" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Rodel, Onadeko, Barej & Sandberger, 2012" ; dwc:species "rainerguentheri" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Rodel & Ernst, 2002" ; dwc:authorityName "Rodel & Ernst" ; dwc:authorityYear "2002" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Rodel & Ernst, 2002" ; dwc:species "phyllophilus" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Guibe & Lamotte, 1962" ; dwc:authorityName "Guibe & Lamotte" ; dwc:authorityYear "1962" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Guibe & Lamotte, 1962" ; dwc:species "guineensis" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Parker, 1936" ; dwc:authorityName "Parker" ; dwc:authorityYear "1936" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Parker, 1936" ; dwc:species "brongersmai" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Rodel, Boateng, Penner, & Hillers" ; dwc:authorityYear "2009" ; dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Rodel, Boateng, Penner, & Hillers, 2009" ; dwc:species "intermedius" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "species" ; dwc:species "tanoeensis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Amphibia" ; dwc:family "Phrynobatrachidae" ; dwc:genus "Phrynobatrachus" ; dwc:kingdom "Animalia" ; dwc:order "Anura" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Phrynobatrachidae" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:order "Anura" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Amphibia" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:phylum "Chordata" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "FIGURE 3. Dorsal and ventral views of adult Phrynobatrachus tanoeensis sp. nov. male (ZMB 86965) from Ebonloa, southwestern Ghana." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 2. Dorsal and ventral views of Phrynobatrachus tanoeensis sp. nov. from the Tanoé-Ehy Swamp Forest region, south-eastern Ivory Coast; a & b) ZMB 80870, holotype, adult male; c & f) ZMB 86957, paratype, adult male; d) ZMB 86956, paratype, adult male; e) ZMB 86960, paratype, adult male; g) ZMB 86961, paratype, adult female; h) ZMB 86961, paratype, adult female." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE1. Neighbour-joiningtree basedonup to 538bp of the 16SRNA gene.Phrynobatrachus tanoeensissp.nov. (yellow) is sister to a clade of Phrynobatrachus species comprising P.intermedius (green) and P.liberiensis (blue); all three being sister to P.tokba (red).Phrynobatrachus liberiensis comprises two clades, one (showing little support) ranging from southern-central Ivory Coast (Banco National Park) west, through western Ivory Coast, south-eastern Guiana and Liberia to eastern Sierra Leone (Gola Rainforest National Park) (pale blue), and one comprising frogs from the Ankasa and Kakum National Parks, south-western Ghana (dark blue). Nodes supported by bootstrap NJ (above) and ML/PP (below); bootstrap value ¿ 65 and PP values> 0.90 shown. Outgroups (P.cricogaster and P. natalensis) not shown." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 4. Advertisement calls of male Phrynobatrachus tanoeensis sp. nov. from Tanoé-Ehy forest, Dohouan village (A, ZMB 86954) and Nouamou (B, ZMB 86958), and P.liberiensis from Taï National Park (C; male not collected; bandpass filter between 1100‒4500 Hz) and Banco National Park (D, ZMB 86969; bandpass filter between 1100‒4500 Hz), all Ivory Coast (compare Table 3); spectrogram (above) and oscillogram (below)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE5. Habitat of Phrynobatrachus tanoeensis sp. nov.; characterized by swampy area, and the presence of Raphia palms with large canopy gaps; at the type locality the Tanoé-Ehy forest, south-eastern Ivory Coast (a, b) and the inundated forest near Ebonloa village, south-western Ghana (c)." ; fabio:hasRepresentation ; a fabio:Figure . dwc:collectionCode "ZMB" ; dwc:eventDate "2010-07-17" ; dwc:locality "GenBank" ; dwc:recordedBy "T. F. Kpan & Adeba." ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "1828873297" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ZMB 80871" ; dwc:collectionCode "ZMB" ; dwc:eventDate "2010-07-17" ; dwc:locality "GenBank" ; dwc:recordedBy "T. F. Kpan & Adeba." ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1828873298" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "ZMB 86953-86955" ; dwc:collectionCode "ZMB" ; dwc:eventDate "2014-10-01" ; dwc:locality "Dohouan village" ; dwc:recordedBy "T. F. Kpan" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "17" ; dwc:verbatimLatitude "5.197117" ; dwc:verbatimLongitude "-2.8856666" ; trt:gbifOccurrenceId "1828873358" ; trt:httpUri ; a dwc:MaterialCitation . dwc:catalogNumber "GAW009" ; dwc:collectionCode "ZMB" ; dwc:locality "Essiama" ; dwc:recordedBy "C. Ofori-Boateng." ; dwc:verbatimElevation "16" ; dwc:verbatimLatitude "5.033411" ; dwc:verbatimLongitude "-2.5516388" ; trt:gbifOccurrenceId "1828873357" ; trt:httpUri ; a dwc:MaterialCitation .