Gromphas inermis Harold 1869
cites: zenodo.3990451
cites: zenodo.3990455
cites: zenodo.3990461
cites: zenodo.3990443
cites: zenodo.3990447
cites: zenodo.3990449
cites: zenodo.3990459
cites: zenodo.3990445
creator: Cupello, Mario; Vaz-de-Mello, Fernando Z.
augments taxon concept: Gromphas inermis Harold 1869
published in: A new species and the phylogeny of the South American genus Gromphas Brullé, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini)
type: Treatment

A new species and the phylogeny of the South American genus Gromphas Brullé, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini)
end page: 969
journal: Journal of Natural History
pub date: 2015-09-30
start page: 943
volume: 50
creator: Cupello, Mario; Vaz-de-Mello, Fernando Z.
date: 2015
has part: zenodo.3990451
has part: zenodo.3990455
has part: zenodo.3990461
has part: zenodo.3990443
has part: zenodo.3990447
has part: zenodo.3990449
has part: zenodo.3990459
has part: zenodo.3990445
type: JournalArticle

Gromphas inermis Harold 1869
authority: Harold, 1869
authority name: Harold
authority year: 1869
class: Insecta
family: Scarabaeidae
genus: Gromphas
kingdom: Animalia
order: Coleoptera
phylum: Arthropoda
rank: species
scientific name authorship: Harold, 1869
species: inermis
has taxon name: Gromphas inermis
type: TaxonConcept

Gromphas inermis
class: Insecta
family: Scarabaeidae
genus: Gromphas
kingdom: Animalia
order: Coleoptera
phylum: Arthropoda
rank: species
species: inermis
has parent name: Gromphas
type: TaxonName

Gromphas
class: Insecta
family: Scarabaeidae
genus: Gromphas
kingdom: Animalia
order: Coleoptera
phylum: Arthropoda
rank: genus
has parent name: Scarabaeidae
type: TaxonName

Scarabaeidae
class: Insecta
family: Scarabaeidae
kingdom: Animalia
order: Coleoptera
phylum: Arthropoda
rank: family
has parent name: Coleoptera
type: TaxonName

Coleoptera
class: Insecta
kingdom: Animalia
order: Coleoptera
phylum: Arthropoda
rank: order
has parent name: Insecta
type: TaxonName

Insecta
class: Insecta
kingdom: Animalia
phylum: Arthropoda
rank: class
has parent name: Arthropoda
type: TaxonName

Arthropoda
kingdom: Animalia
phylum: Arthropoda
rank: phylum
has parent name: Animalia
type: TaxonName

Animalia
kingdom: Animalia
rank: kingdom
type: TaxonName

zenodo.3990451
description: Figure 5. Updated distribution of the six species of Gromphas.
type: Figure

zenodo.3990455
description: Figure 6. Phylogeny of Gromphas. Above: Cladogram showing the phylogenetic relationship of the species of Gromphas and the evolution of their characters. Black circles indicate uncontroverted synapormorphies; white circles with black margin, controverted synapomorphies; red circles, ambiguous uncontroverted synapomorphies; and white circles with red margin, ambiguous controverted synapomorphies. Below: Support values given for each clade are decay index (above branch) and bootstrap (below branch).
type: Figure

zenodo.3990461
description: Figure 8. Lateral view of male protibia, ornamentation of ventral carina. (A) Gromphas inermis: a row of tubercles (indicated by red arrow), a synapomorphy of clade B of Gromphas. (B) Oruscatus davus: a single large tooth. (C) Bolbites onitoides: a large central tooth with minor lateral teeth.
type: Figure

zenodo.3990443
description: Figure 1. Gromphas jardim sp. nov. (A–D) Holotype. (A) Dorsal view. (B) Lateral view. (C) Aedeagus. From left to right, dorsal, lateral and ventral views. (D) Labels. (E) Female paratype from Bolivia. (F) Dorsal view of head (holotype). (G) Frontal view of cephalic projection (male paratype).
type: Figure

zenodo.3990447
description: Figure 3. Lateral view of pronotum. (A) Gromphas jardim sp. nov. (B) Gromphas amazonica. Note that the granulation extends much more posteriorly in G. jardim, reaching the pronotal posterior margin, than in G. amazonica, wherein the granulation is absent or rudimentary in posterolateral region after lateral fossa.
type: Figure

zenodo.3990449
description: Figure 4. Male paratype of Gromphas jardim sp. nov. and its labels (braces ‘{’ joining two sides of a same label). We believe that the French zoologist and explorer Alcide d’Orbigny was the collector of this specimen in 1832 (see the text).
type: Figure

zenodo.3990459
description: Figure 7. Metepisterna. (A) Gromphas jardim sp. nov.: gromphas is unique among the Phanaeini by not showing any trace of a metepisternal tab (MTab) covering the epipleural margin, a synapomorphy of this genus. (B) Oruscatus davus: Both species of Oruscatus have a small but evident MTab (indicated by red arrow). (C) Bolbites onitoides: The MTab of Bolbites is strongly developed and the dorsal margin of metepisternum is highly curved. This is the same form seen in the rest of Phanaeina and is a synapomorphy of this taxon. (D) Dichotomius fissus (Harold, 1867): MTab in a non-Phanaeini. This structure is present also in a series of other Coprini, as well as in at least some Oniticellini and Onitini.
type: Figure

zenodo.3990445
description: Figure 2. Protibia of Gromphas jardim sp. nov. (A, B) Male protibia: (A) Ventral view, (B) Dorsal view. Red arrow indicates apical tubercle modified in a tapered spur. (C) Female protibia. Note that the apical tuft of setae is much denser and longer in male than in female.
type: Figure

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locality: Rio San Javier
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recorded by: Pelotas
gbif occurrence id: 2831865425
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A43142DFC6D3545FC03
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collection code: BMNH
gbif occurrence id: 2831865416
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A4314FBFC6D36F4FC61
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collection code: BMNH, OUMNH
event date: 1820-03
locality: SANTA CATARINA
recorded by: Hope-Westwood
gbif occurrence id: 2831865409
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A43160FFC4F342BFC44
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collection code: BMNH, MACN
locality: PARAGUAY
gbif occurrence id: 2831865459
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A43153BFC2A37F1FBAB
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collection code: BMNH, MACN
locality: PARAGUAY
recorded by: Richter
gbif occurrence id: 2831865358
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A4316F7FC153486FBAB
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collection code: MLPA
gbif occurrence id: 2831865453
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A431581FC1535B6FBAB
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collection code: MLPA
gbif occurrence id: 2831865324
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A4314B1FC1535D4FBAB
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collection code: MLPA
gbif occurrence id: 2831865433
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A43135AFC1537EBFB89
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event date: 1922-09
locality: Asuncion
gbif occurrence id: 2831865477
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A4316FCFBF736F7FBEC
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event date: 1922-09
locality: Asuncion
gbif occurrence id: 2831865335
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A4317F1FBD23715FBEC
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collection code: BMNH
gbif occurrence id: 2831865420
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A431685FBD23470FBEC
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locality: Villarrica
gbif occurrence id: 2831865351
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A431585FBD2322BFBEC
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locality: Villarrica
recorded by: Richter
gbif occurrence id: 2831865424
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A43132CFBD236CAFBD3
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collection code: MLPA
gbif occurrence id: 2831865400
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A4317CDFBBD345DFBD3
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locality: Sapucai
gbif occurrence id: 2831865332
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A43156BFBBD35C9FBD3
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locality: Sapucai
recorded by: Foster
gbif occurrence id: 2831865472
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A4314C1FBBD3243FBD3
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collection code: BMNH
gbif occurrence id: 2831865471
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A431344FBBD376CFB31
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collection code: BMNH, MACN
locality: URUGUAY
recorded by: Hermann Burmeister
gbif occurrence id: 2831865446
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A43167DFB9F3493FB14
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collection code: MACN
locality: Atlantida
gbif occurrence id: 2831865427
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A4315A3FB7A3732FB7B
type: MaterialCitation

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collection code: MACN
locality: COLONIA
recorded by: Riachuelo
gbif occurrence id: 2831865392
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A431640FB25328EFB7B
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collection code: MACN
locality: COLONIA
recorded by: Stevenin
gbif occurrence id: 2831865445
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A431389FB253739FB59
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collection code: MACN
gbif occurrence id: 2831865325
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A43163EFB07342BFB59
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collection code: BMNH, MACN, MLPA
event date: 1929-12-20
locality: Penarol
recorded by: Penarol
gbif occurrence id: 2831865457
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A431531FB0736FAFAC1
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collection code: MACN
locality: Bruch-Waiser
recorded by: Carlos Bruch & Parana
gbif occurrence id: 2831865385
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A431608FAAF374EFA24
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locality: Nov.
recorded by: Brazil
gbif occurrence id: 2831865315
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A431646FA8A349FFA24
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collection code: MACN
recorded by: Hermann Burmeister
gbif occurrence id: 2831865388
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A4315B0FA8A36D6FA0B
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collection code: BMNH
locality: Specimens
recorded by: Frey
gbif occurrence id: 2831865336
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A4317F9FA753780FA69
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collection code: MACN, OUMNH
locality: No
recorded by: Carlos Bruch & Hermann Burmeister & Westwood & Gory
gbif occurrence id: 2831865397
http uri: https://treatment.plazi.org/id/039B9101BC582A5A16D9FBF735D0FBC3#3B5A2A4ABC5A2A431696FA57351EF9B3
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