@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , ; dc:creator "Kompier, Tom; Karube, Haruki" ; dc:title "Heliogomphus bidentatus Kompier & Karube 2019, sp. nov." ; dwc:basisOfRecord , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; trt:citesTaxonName ; trt:definesTaxonConcept ; trt:deprecates , , , , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "120" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:pubDate "2019-02-12" ; bibo:startPage "113" ; bibo:volume "4555" ; dc:creator "Kompier, Tom; Karube, Haruki" ; dc:date "2019" ; dc:title "A new species of Heliogomphus from Vietnam (Odonata: Gomphidae)" ; fabio:hasPart , , , , ; a fabio:JournalArticle . dwc:authority "Kompier & Karube, 2019" ; dwc:authorityName "Kompier & Karube" ; dwc:authorityYear "2019" ; dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Kompier & Karube, 2019" ; dwc:species "bidentatus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Fraser, 1942" ; dwc:authorityName "Fraser" ; dwc:authorityYear "1942" ; dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Fraser, 1942" ; dwc:species "lieftincki" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Ris" ; dwc:baseAuthorityYear "1912" ; dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Ris, 1912)" ; dwc:species "scorpio" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Lieftinck, 1961" ; dwc:authorityName "Lieftinck" ; dwc:authorityYear "1961" ; dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Lieftinck, 1961" ; dwc:species "olivaceus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Lieftinck, 1964" ; dwc:authorityName "Lieftinck" ; dwc:authorityYear "1964" ; dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Lieftinck, 1964" ; dwc:species "borneensis" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Karube, 2004" ; dwc:authorityName "Karube" ; dwc:authorityYear "2004" ; dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Karube, 2004" ; dwc:species "chaoi" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Laidlaw" ; dwc:authorityYear "1902" ; dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Laidlaw, 1902" ; dwc:species "kelantanensis" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "bidentatus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "lieftincki" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "scorpio" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Pneumoridae" ; dwc:genus "Vesica" ; dwc:kingdom "Animalia" ; dwc:order "Orthoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "spermalis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Pneumoridae" ; dwc:genus "Vesica" ; dwc:kingdom "Animalia" ; dwc:order "Orthoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Pneumoridae" ; dwc:kingdom "Animalia" ; dwc:order "Orthoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Orthoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "olivaceus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "borneensis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "chaoi" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Gomphidae" ; dwc:genus "Heliogomphus" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "kelantanensis" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIGURE 1. Habitus of Heliogomphus bidentatus [a–b, d–e] and H. scorpio [c, f]. (a) male, 5-V-2016, Quang Tri Prov.; (b) paratype male, 11-V-2014, Nam Bung, Yen Bai Prov.; (c) male, 17-V-2014, Tam Dao NP, Vinh Phuc Prov.; (d) paratype female, 15-VII-2017, Pia Oac NR, Cao Bang Prov.; (e) paratype female (used for description), 16-V-2015, Nam Bung, Yen Bai Prov.; (f) female, 15-VI-2015, Thai Nguyen Prov.." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 4. Vesica spermalis (a) and hamuli (b) of holotype Heliogomphus bidentatus in lateral view." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 3. Comparison of male caudal appendages of holotype Heliogomphus bidentatus [a–c] and H. scorpio from Guangdong Prov., China (collection Karube) [d–f] in dorsal view (a, d), lateral view (b, e) and ventral view of epiproct (c, f)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 2. Comparison of male caudal appendages in dorsal view of (a) Heliogomphus bidentatus (11-V-2014, Nam Bung, Yen Bai Prov.) and (b) H. scorpio (17-V-2014, Tam Dao NP, Vinh Phuc Prov.)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 5. Occipital area of (a) female Heliogomphus bidentatus (paratype, 16-V-2015, Nam Bung, Yen Bai Prov.) and (b) female H. scorpio (30-V-2015, Xuan Son NP, Phu Tho Prov.)." ; fabio:hasRepresentation ; a fabio:Figure . dwc:eventDate "1994-05-02" ; dwc:locality "Vinh Phuc" ; dwc:recordedBy "Tam Dao NP" ; dwc:typeStatus "holotype" ; trt:gbifOccurrenceId "2234218913" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "1993-05-19" ; dwc:locality "Vinh Phuc" ; dwc:recordedBy "Tam Dao NP" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "2234218883" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "1997-05-31" ; dwc:locality "Tam Dao" ; dwc:recordedBy "Y. Arita" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "2234218912" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "1998-05-02" ; dwc:locality "Prov." ; dwc:recordedBy "Pia Oac NR & K. Matsumoto" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "2234218845" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2014-05-11" ; dwc:locality "Prov." ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "2234218827" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2014-05-20" ; dwc:locality "Tu Le" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "2234218875" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2014-07-09" ; dwc:locality "Prov." ; dwc:recordedBy "Pia Oac NR" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "2234218873" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2015-05-03" ; dwc:locality "Mt." ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "2234218887" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2015-05-16" ; dwc:locality "Prov." ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "21.74" ; dwc:verbatimLongitude "104.277" ; trt:gbifOccurrenceId "2234218829" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V, T" ; dwc:eventDate "2015-05-19" ; dwc:locality "Prov." ; dwc:recordedBy "Phan" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "566" ; trt:gbifOccurrenceId "2234218833" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2016-05-19" ; dwc:locality "Prov." ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "16.09" ; dwc:verbatimLongitude "107.464" ; trt:gbifOccurrenceId "2234218821" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2017-07-15" ; dwc:locality "Prov." ; dwc:recordedBy "Pia Oac NR" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "2234218955" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2018-06-08" ; dwc:locality "Prov." ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "2234218854" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2018-06-10" ; dwc:locality "Prov." ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "21.752" ; dwc:verbatimLongitude "104.276" ; trt:gbifOccurrenceId "2234218820" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2014-05-11" ; dwc:locality "Prov." ; dwc:recordedBy "Nam Bung" ; trt:gbifOccurrenceId "2234218881" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2014-06-15" ; dwc:locality "Prov" ; dwc:recordedBy "Tam Dao NP" ; trt:gbifOccurrenceId "2234218817" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2014-07-04" ; dwc:locality "Prov." ; trt:gbifOccurrenceId "2234218835" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2014-07-09" ; dwc:locality "Prov." ; dwc:recordedBy "Pia Oac NR" ; trt:gbifOccurrenceId "2234218909" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2015-04-18" ; dwc:locality "Prov." ; dwc:recordedBy "Xuan Son NP" ; trt:gbifOccurrenceId "2234218910" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2015-05-01" ; dwc:locality "Prov." ; dwc:recordedBy "Pia Oac NR" ; trt:gbifOccurrenceId "2234218886" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2015-05-11" ; dwc:locality "Prov." ; dwc:recordedBy "Pu Luong NR" ; trt:gbifOccurrenceId "2234218900" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2015-05-14" ; dwc:locality "Prov." ; trt:gbifOccurrenceId "2234218922" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2015-05-16" ; dwc:locality "Prov." ; trt:gbifOccurrenceId "2234218959" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2015-05-30" ; dwc:locality "Prov." ; dwc:recordedBy "Xuan Son NP" ; trt:gbifOccurrenceId "2234218870" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2015-05-31" ; dwc:locality "Prov." ; dwc:recordedBy "Pia Oac NR" ; trt:gbifOccurrenceId "2234218844" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2015-06-14" ; dwc:locality "Prov." ; trt:gbifOccurrenceId "2234218868" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2015-06-17" ; dwc:locality "Prov." ; dwc:recordedBy "Pia Oac NR" ; trt:gbifOccurrenceId "2234218931" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2015-06-18" ; dwc:locality "Prov." ; dwc:recordedBy "Pia Oac NR" ; trt:gbifOccurrenceId "2234218840" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2015-06-29" ; dwc:locality "Prov." ; dwc:recordedBy "Xuan Son NP" ; trt:gbifOccurrenceId "2234218890" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2016-05-05" ; dwc:locality "Prov." ; dwc:recordedBy "Highway West" ; trt:gbifOccurrenceId "2234218808" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "V" ; dwc:eventDate "2016-05-19" ; dwc:locality "Prov." ; dwc:recordedBy "Highway" ; trt:gbifOccurrenceId "2234218884" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2016-06-03" ; dwc:locality "Prov." ; dwc:recordedBy "Pia Oac NR" ; trt:gbifOccurrenceId "2234218876" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2016-06-04" ; dwc:locality "Prov." ; dwc:recordedBy "Tam Dao NP" ; trt:gbifOccurrenceId "2234218850" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2018-06-06" ; dwc:locality "Ha Noi" ; dwc:recordedBy "Ba Vi NP" ; trt:gbifOccurrenceId "2234218899" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2018-06-08" ; dwc:locality "Prov." ; dwc:recordedBy "Xuan Son NP" ; trt:gbifOccurrenceId "2234218898" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "VI" ; dwc:eventDate "2018-06-09" ; dwc:locality "Prov." ; dwc:recordedBy "Xuan Son NP" ; trt:gbifOccurrenceId "2234218939" ; trt:httpUri ; a dwc:MaterialCitation .