@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , ; dc:creator "Génier, François; Moretto, Philippe" ; dc:title "Digitonthophagus sahelicus Moretto, new species" ; dwc:basisOfRecord , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; trt:definesTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "110" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:startPage "1" ; bibo:volume "4248" ; dc:creator "Génier, François; Moretto, Philippe" ; dc:date "2017" ; dc:title "Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae)" ; fabio:hasPart , , , , , ; a fabio:JournalArticle . dwc:authority "Moretto" ; dwc:authorityName "Moretto" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Digitonthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Moretto" ; dwc:species "sahelicus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Digitonthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "sahelicus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Digitonthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "FIGURES 7 – 10. Male dorsal habitus. 7. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 8. D. uks, holotype; 9. D. sahelicus, holotype; 10. D. catta, alloreferent (Tamil Nadu State, India)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 23 – 30. 23, 25, 27, 29. Male head, dorsal view; 24, 26, 28, 30. Male head, frontal view. 23 – 24. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 25 – 26. D. uks, holotype; 27 – 28. D. sahelicus, holotype; 29 – 30. D. catta, alloreferent (Tamil Nadu State, India)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 55 – 62. Male pronotum, dorsal view. 55. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 56. D. uks, holotype; 57. D. sahelicus, holotype; 58. D. catta, alloreferent (Tamil Nadu State, India); 59. D. eucatta, holotype; 60. D. falciger, holotype; 61. D. biflagellatus, holotype; 62. D. gazella, non type (Central Province, Zambia)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 71 – 78. Female head, dorsal view. 71. Digitonthophagus bonasus, non type (Tamil Nadu State, India); 72. D. uks, allotype; 73. D. sahelicus, allotype; 74. D. catta, non type (India); 75. D. eucatta, allotype; 76. D. falciger, allotype; 77. D. biflagellatus, allotype; 78. D. gazella, non type (Antisiranana Province, Madagascar)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 119 – 134. Parameres, dorsal view. 119. D. bonasus, alloreferent (Tamil Nadu State, India); 120. Digitonthophagus uks, holotype; 121. D. sahelicus, holotype; 122. D. catta, alloreferent (West Bengal, India); 123. D. eucatta, holotype; 124. D. falciger, holotype; 125. D. biflagellatus, holotype; 126. D. gazella, non type (Antisiranana Province, Madagascar); 127. D. petillus, holotype; 128. D. viridicollis, holotype; 129. D. aksumensis, holotype; 130. D. dilatatus, holotype; 131. D. fimator, holotype; 132. D. lusinganus, lectotype; 133. D. ulcerosus, holotype 134. D. namaquensis, holotype." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 135 – 146. Internal sac sclerites (FLP, A + SA). 135, 138, 141, 144. Dorsal view; 136, 139, 142, 145. Ventral view. 137, 140, 143, 146. Right lateral view. 135 – 137. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 138 – 140. D. uks, holotype; 141 – 143. D. sahelicus, holotype; 144 – 146. D. catta, alloreferent." ; fabio:hasRepresentation ; a fabio:Figure . dwc:locality "Lorum" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "300" ; dwc:verbatimLatitude "13.895278" ; dwc:verbatimLongitude "-1.8736111" ; trt:gbifOccurrenceId "1455385747" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC, LORUM" ; dwc:eventDate "2006" ; dwc:locality "Primary" ; dwc:recordedBy "F. & S. Genier & P. Moretto" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "300" ; dwc:verbatimLatitude "13.895278" ; trt:gbifOccurrenceId "1455385746" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2002-08" ; dwc:locality "Bobo-Dioulasso" ; dwc:verbatimLatitude "11.2" ; dwc:verbatimLongitude "-4.3" ; trt:gbifOccurrenceId "1455385687" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "PMOC" ; dwc:eventDate "2012-08-07" ; dwc:locality "Gorum-Gorum" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "271" ; dwc:verbatimLatitude "14.159111" ; dwc:verbatimLongitude "0.022333335" ; trt:gbifOccurrenceId "1455385712" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "PMOC" ; dwc:eventDate "2013" ; dwc:locality "Dori" ; dwc:recordedBy "P. Moretto" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "283" ; dwc:verbatimLatitude "14.034584" ; dwc:verbatimLongitude "-0.05291667" ; trt:gbifOccurrenceId "1455385752" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "PMOC" ; dwc:eventDate "2013-08-30" ; dwc:locality "Dori" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "283" ; dwc:verbatimLatitude "14.034584" ; dwc:verbatimLongitude "-0.05291667" ; trt:gbifOccurrenceId "1455385764" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "JFJC" ; dwc:eventDate "2007-04" ; dwc:locality "Dori" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "14.033334" ; dwc:verbatimLongitude "-0.016666668" ; trt:gbifOccurrenceId "1455385769" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "PMOC" ; dwc:eventDate "2012-08-13" ; dwc:locality "Gorum-Gorum" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "264" ; dwc:verbatimLatitude "14.377833" ; dwc:verbatimLongitude "0.09658334" ; trt:gbifOccurrenceId "1455385740" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "PMOC" ; dwc:eventDate "2013-08-24" ; dwc:locality "Dori" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "14.065473" ; dwc:verbatimLongitude "-0.050944444" ; trt:gbifOccurrenceId "1455385770" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FGIC" ; dwc:eventDate "2006" ; dwc:locality "Toulfe" ; dwc:recordedBy "F. & S. Genier" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "300" ; dwc:verbatimLatitude "13.895278" ; dwc:verbatimLongitude "-1.8736111" ; trt:gbifOccurrenceId "1455385725" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FGIC" ; dwc:eventDate "2006" ; dwc:locality "Toulfe" ; dwc:recordedBy "F. & S. Genier" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "300" ; dwc:verbatimLatitude "13.895278" ; dwc:verbatimLongitude "-1.8736111" ; trt:gbifOccurrenceId "1455385757" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FGIC" ; dwc:eventDate "2006" ; dwc:locality "Toulfe" ; dwc:recordedBy "F. & S. Genier" ; dwc:typeStatus "paratype" ; dwc:verbatimElevation "300" ; dwc:verbatimLatitude "13.895278" ; dwc:verbatimLongitude "-1.8736111" ; trt:gbifOccurrenceId "1455385780" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC, FGIC" ; dwc:eventDate "2006" ; dwc:locality "Toulfe" ; dwc:recordedBy "F. & S. Genier" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "300" ; dwc:verbatimLatitude "13.895278" ; dwc:verbatimLongitude "-1.8736111" ; trt:gbifOccurrenceId "1455385760" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FGIC" ; dwc:eventDate "2006" ; dwc:locality "Toulfe" ; dwc:recordedBy "F. & S. Genier" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "300" ; dwc:verbatimLatitude "13.895278" ; dwc:verbatimLongitude "-1.8736111" ; trt:gbifOccurrenceId "1455385719" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FGIC" ; dwc:eventDate "2006" ; dwc:locality "Boulieselo" ; dwc:recordedBy "Foret de Nazinga & F. & S. Genier" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "310" ; dwc:verbatimLatitude "11.197223" ; dwc:verbatimLongitude "-1.5858333" ; trt:gbifOccurrenceId "1455385761" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "EXTREME, PMOC" ; dwc:eventDate "2006-08-18" ; dwc:locality "Waza National Park" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "11.25" ; dwc:verbatimLongitude "14.65" ; trt:gbifOccurrenceId "1455385766" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:eventDate "1935-04-09" ; dwc:locality "Fada" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "17.183332" ; dwc:verbatimLongitude "21.583334" ; trt:gbifOccurrenceId "1455385790" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "PMOC" ; dwc:eventDate "2006-08-11" ; dwc:locality "Sitje" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "12.883333" ; dwc:verbatimLongitude "14.866667" ; trt:gbifOccurrenceId "1455385727" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "1935-07-19" ; dwc:locality "Ziguei" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "14.716666" ; dwc:verbatimLongitude "15.783334" ; trt:gbifOccurrenceId "1455385729" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "DJIBOUTI" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455385755" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IRSNB" ; dwc:locality "Djibouti" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "11.583333" ; dwc:verbatimLongitude "43.15" ; trt:gbifOccurrenceId "1455385742" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "PMOC" ; dwc:eventDate "2003-07" ; dwc:locality "Hagaz" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "15.7" ; dwc:verbatimLongitude "38.266666" ; trt:gbifOccurrenceId "1455385777" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FGIC, PMOC" ; dwc:eventDate "2003-07" ; dwc:locality "Akordat" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "15.55" ; dwc:verbatimLongitude "38.016666" ; trt:gbifOccurrenceId "1455385734" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FETC" ; dwc:eventDate "2002-07" ; dwc:locality "Nefasit" ; dwc:recordedBy "Czeppel" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "15.333333" ; dwc:verbatimLongitude "39.066666" ; trt:gbifOccurrenceId "1455385739" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "SEGOU, MNHN" ; dwc:locality "Sansanding" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "13.721666" ; dwc:verbatimLongitude "-6.0030556" ; trt:gbifOccurrenceId "1455385730" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "IRSNB" ; dwc:locality "Trarza" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455385728" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "PMOC" ; dwc:eventDate "1968-09-24" ; dwc:locality "Atar" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "20.516666" ; dwc:verbatimLongitude "-13.05" ; trt:gbifOccurrenceId "1455385758" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:eventDate "1937-10" ; dwc:locality "Oued Akjoujt" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "19.75" ; dwc:verbatimLongitude "-14.383333" ; trt:gbifOccurrenceId "1455385765" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "PMOC" ; dwc:eventDate "2006-08-15" ; dwc:locality "Nouakchoot" ; dwc:recordedBy "Famille" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "18.083334" ; dwc:verbatimLongitude "-15.983334" ; trt:gbifOccurrenceId "1455385721" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OMOC" ; dwc:eventDate "1941-10-20" ; dwc:locality "Tachoot" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "15.433333" ; dwc:verbatimLongitude "-12.266666" ; trt:gbifOccurrenceId "1455385718" ; trt:httpUri ; a dwc:MaterialCitation . dwc:locality "AGADEZ" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "1455385775" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "Agadez" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "16.966667" ; dwc:verbatimLongitude "7.9833336" ; trt:gbifOccurrenceId "1455385753" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:eventDate "1947-08-02" ; dwc:locality "Agadez" ; dwc:recordedBy "L. 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Chopard" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "16.966667" ; dwc:verbatimLongitude "7.9833336" ; trt:gbifOccurrenceId "1455385802" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "OMOC" ; dwc:eventDate "1989-08-20" ; dwc:locality "Agadez" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "16.966667" ; dwc:verbatimLongitude "7.9833336" ; trt:gbifOccurrenceId "1455385785" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "PMOC" ; dwc:eventDate "1977-09-15" ; dwc:locality "Arlit" ; dwc:typeStatus "paratype" ; dwc:verbatimLatitude "18.733334" ; dwc:verbatimLongitude "7.383333" ; trt:gbifOccurrenceId "1455385741" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:eventDate "1947-08-13" ; dwc:locality "Air" ; dwc:recordedBy "L. 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