@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , , , ; dc:creator "Génier, François; Moretto, Philippe" ; dc:title "Digitonthophagus gazella Fabricius 1787" ; dwc:basisOfRecord , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; trt:augmentsTaxonConcept ; trt:citesTaxonName , , , , , , , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "110" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:startPage "1" ; bibo:volume "4248" ; dc:creator "Génier, François; Moretto, Philippe" ; dc:date "2017" ; dc:title "Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae)" ; fabio:hasPart , , , , , , ; a fabio:JournalArticle . dwc:authority "Fabricius, 1787" ; dwc:authorityName "Fabricius" ; dwc:authorityYear "1787" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Digitonthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Fabricius, 1787" ; dwc:species "gazella" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Gazella Fabricius, 1787b: 377" ; dwc:authorityName "Gazella Fabricius" ; dwc:authorityPageNumber "377" ; dwc:authorityYear "1787" ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Scarabaeus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Gazella Fabricius, 1787" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Digitonthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "gazella" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Digitonthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Scarabaeus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Scarabaeus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "gazella" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Copris" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "bonasus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Aphodiidae" ; dwc:genus "Ontophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "gazella" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Aphodiidae" ; dwc:genus "Ontophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Aphodiidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . # Warning: abbreviated subGenus "O." ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "gazella" ; dwc:subGenus "O.", "Digitonthophagus" ; trt:hasParentName , , ; a dwcFP:TaxonName . # Warning: abbreviated subGenus "O." ; dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:subGenus "O." ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:subGenus "Digitonthophagus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Scarabaeidae" ; dwc:genus "Onthophagus" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "dorcas" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIGURES 11 – 14. Male dorsal habitus. 11. D. eucatta, holotype; 12. D. falciger, holotype; 13. Digitonthophagus biflagellatus, holotype; 14. D. gazella, non type (Central Province, Zambia)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 31 – 38. 31, 33, 35, 37. Male head, dorsal view; 32, 34, 36, 38. Male head, frontal view. 31 – 32. Digitonthophagus eucatta, alloreferent; 33 – 34. D. falciger, holotype; 35 – 36. D. biflagellatus, holotype; 37 – 38. D. gazella, non type (Central Province, Zambia)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 55 – 62. Male pronotum, dorsal view. 55. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 56. D. uks, holotype; 57. D. sahelicus, holotype; 58. D. catta, alloreferent (Tamil Nadu State, India); 59. D. eucatta, holotype; 60. D. falciger, holotype; 61. D. biflagellatus, holotype; 62. D. gazella, non type (Central Province, Zambia)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 71 – 78. Female head, dorsal view. 71. Digitonthophagus bonasus, non type (Tamil Nadu State, India); 72. D. uks, allotype; 73. D. sahelicus, allotype; 74. D. catta, non type (India); 75. D. eucatta, allotype; 76. D. falciger, allotype; 77. D. biflagellatus, allotype; 78. D. gazella, non type (Antisiranana Province, Madagascar)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 147 – 158. Internal sac sclerites (FLP, A + SA). 147, 150, 153, 156. Dorsal view; 148, 151, 154, 157. Ventral view. 149, 152, 155, 158. Right lateral view. 147 – 149. Digitonthophagus eucatta, alloreferent (Tamil Nadu State, India); 150 – 152. D. falciger, holotype; 153 – 155. D. biflagellatus, holotype; 156 – 158. D. gazella, non type (Antisiranana Province, Madagascar)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 183 – 190. Digitonthophagus bonasus, female head, frontal view (Phongsaly Province, Laos); 184. D. aksumensis, FLP sclerite, right lateral view; 185. D. gazella, right lateral fold, ventral view; 186. D. viridicollis, right lateral fold, ventral view; 187. D. eucatta, hypomeron, ventral view, A. anterior hypomeral carina, B. anterior hypomeral depression; D. gazella, hypomeron, ventral view, A. anterior hypomeral carina, B. anterior hypomeral depression; 189. D. namaquensis, vertex dorsal view; 190. D. gazella, vertex dorsal view." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURES 119 – 134. Parameres, dorsal view. 119. D. bonasus, alloreferent (Tamil Nadu State, India); 120. Digitonthophagus uks, holotype; 121. D. sahelicus, holotype; 122. D. catta, alloreferent (West Bengal, India); 123. D. eucatta, holotype; 124. D. falciger, holotype; 125. D. biflagellatus, holotype; 126. D. gazella, non type (Antisiranana Province, Madagascar); 127. D. petillus, holotype; 128. D. viridicollis, holotype; 129. D. aksumensis, holotype; 130. D. dilatatus, holotype; 131. D. fimator, holotype; 132. D. lusinganus, lectotype; 133. D. ulcerosus, holotype 134. D. namaquensis, holotype." ; fabio:hasRepresentation ; a fabio:Figure . dwc:collectionCode "MNHN" ; dwc:eventDate "1914-03" ; dwc:locality "Menongue" ; dwc:recordedBy "Mission" ; dwc:verbatimLatitude "-14.65" ; dwc:verbatimLongitude "17.683332" ; trt:gbifOccurrenceId "1455384743" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FGIC, GWPC" ; dwc:eventDate "2011-11-05" ; dwc:locality "Caconda" ; dwc:verbatimLatitude "-13.833333" ; dwc:verbatimLongitude "15.1" ; trt:gbifOccurrenceId "1455384714" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "GWPC" ; dwc:eventDate "2011-11-04" ; dwc:locality "Cacula" ; dwc:verbatimLatitude "-14.583333" ; dwc:verbatimLongitude "14.05" ; trt:gbifOccurrenceId "1455384769" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "FGIC, GWPC" ; dwc:eventDate "2013-11-26" ; dwc:locality "Quipungo" ; dwc:verbatimLatitude "-14.800333" ; dwc:verbatimLongitude "14.255333" ; trt:gbifOccurrenceId "1455384748" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "GWPC" ; dwc:eventDate "2011-11-13" ; dwc:locality "Cacula" ; dwc:verbatimLatitude "-14.683333" ; dwc:verbatimLongitude "13.9" ; trt:gbifOccurrenceId "1455384785" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:eventDate "1914-12" ; dwc:locality "Luceque" ; dwc:recordedBy "Mission" ; dwc:verbatimLatitude "-14.65" ; dwc:verbatimLongitude "15.066667" ; trt:gbifOccurrenceId "1455384747" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:locality "Capelongo-Dongo" ; dwc:recordedBy "Mission" ; dwc:verbatimLatitude "-14.516666" ; dwc:verbatimLongitude "16.016666" ; trt:gbifOccurrenceId "1455384762" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "MNHN" ; dwc:eventDate "1914-12" ; dwc:locality "Capelongo-Dongo" ; dwc:recordedBy "Mission" ; dwc:verbatimLatitude "-14.516666" ; dwc:verbatimLongitude "16.016666" ; trt:gbifOccurrenceId "1455384761" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:eventDate "1988-12-05" ; dwc:locality "Warrumbungle National Park" ; dwc:verbatimLatitude "-31.276945" ; dwc:verbatimLongitude "148.9975" ; trt:gbifOccurrenceId "1455384763" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:eventDate "1988-04-12" ; dwc:locality "Gilgandra" ; dwc:verbatimLatitude "-31.716667" ; dwc:verbatimLongitude "148.66667" ; trt:gbifOccurrenceId "1455384770" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:eventDate "1977-12-05" ; dwc:locality "Narrabri" ; dwc:recordedBy "H.G. Garcia" ; dwc:verbatimLatitude "-30.333334" ; dwc:verbatimLongitude "149.78334" ; trt:gbifOccurrenceId "1455384764" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CSIRO, CMNC" ; dwc:eventDate "1993-12-01" ; dwc:locality "Darwin" ; dwc:recordedBy "S. & J. Peck" ; dwc:verbatimLatitude "-12.411666" ; dwc:verbatimLongitude "130.92195" ; trt:gbifOccurrenceId "1455384765" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CSIRO, CMNC" ; dwc:eventDate "1994-01-05" ; dwc:locality "Darwin" ; dwc:recordedBy "S. & J. Peck" ; dwc:verbatimLatitude "-12.411666" ; dwc:verbatimLongitude "130.92195" ; trt:gbifOccurrenceId "1455384768" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CSIRO, CMNC" ; dwc:eventDate "1994-01-09" ; dwc:locality "Darwin" ; dwc:recordedBy "S. & J. Peck" ; dwc:verbatimLatitude "-12.411666" ; dwc:verbatimLongitude "130.92195" ; trt:gbifOccurrenceId "1455384775" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:eventDate "1993-12-09" ; dwc:locality "Mataranka" ; dwc:recordedBy "S. & J. Peck" ; dwc:verbatimLatitude "-14.966666" ; dwc:verbatimLongitude "133.18333" ; trt:gbifOccurrenceId "1455384760" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:eventDate "1993-12-09" ; dwc:locality "Mataranka" ; dwc:recordedBy "S. & J. Peck" ; dwc:verbatimLatitude "-14.966666" ; dwc:verbatimLongitude "133.18333" ; trt:gbifOccurrenceId "1455384753" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:eventDate "1993-12-22" ; dwc:locality "Kakadu National Park" ; dwc:recordedBy "S. & J. Peck" ; dwc:verbatimLatitude "-12.679444" ; dwc:verbatimLongitude "132.48584" ; trt:gbifOccurrenceId "1455384750" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:eventDate "1989-11-04" ; dwc:locality "Magela Creek" ; dwc:recordedBy "L. 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trt:gbifOccurrenceId "1455384754" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CEMT" ; dwc:eventDate "1993-02" ; dwc:locality "Lynd" ; dwc:verbatimLatitude "-18.6" ; dwc:verbatimLongitude "144.71666" ; trt:gbifOccurrenceId "1455384776" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:eventDate "1997-01-28" ; dwc:locality "Mareeba" ; dwc:verbatimLatitude "-16.916666" ; dwc:verbatimLongitude "145.41667" ; trt:gbifOccurrenceId "1455384752" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:eventDate "1986-12-13" ; dwc:locality "Julatten" ; dwc:verbatimLatitude "-16.55" ; dwc:verbatimLongitude "145.38333" ; trt:gbifOccurrenceId "1455384744" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:eventDate "1975-02-07" ; dwc:locality "Atherton" ; dwc:verbatimLatitude "-17.266666" ; dwc:verbatimLongitude "145.48334" ; trt:gbifOccurrenceId "1455384767" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; 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dwc:verbatimLongitude "145.76666" ; trt:gbifOccurrenceId "1455384772" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "1975-01-30" ; dwc:locality "Cairns" ; dwc:verbatimLatitude "-16.916666" ; dwc:verbatimLongitude "145.76666" ; trt:gbifOccurrenceId "1455384788" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:locality "A" ; trt:gbifOccurrenceId "1455384782" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:eventDate "1975-02-03" ; dwc:locality "Cairns" ; dwc:verbatimLatitude "-16.916666" ; dwc:verbatimLongitude "145.76666" ; trt:gbifOccurrenceId "1455384783" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:eventDate "1975-02-05" ; dwc:locality "Cairns" ; dwc:verbatimLatitude "-16.916666" ; dwc:verbatimLongitude "145.76666" ; trt:gbifOccurrenceId "1455384795" ; trt:httpUri ; a dwc:MaterialCitation . dwc:collectionCode "CMNC" ; dwc:eventDate "1988-12-16" ; dwc:locality "Charters Towers" ; dwc:verbatimLatitude "-20.083334" ; 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