@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , ; dc:creator "Pinto-Da-Rocha, Ricardo; Cabra-García, Jimmy" ; dc:title "Tityus forcipula" ; trt:augmentsTaxonConcept ; trt:deprecates , , , , , , , , , , , , , , , , , , , , ; trt:publishedIn ; a trt:Treatment . bibo:endPage "186" ; bibo:issue "2" ; bibo:journal "Zootaxa" ; bibo:pubDate "2022-06-21" ; bibo:startPage "151" ; bibo:volume "5155" ; dc:creator "Pinto-Da-Rocha, Ricardo; Cabra-García, Jimmy" ; dc:date "2022" ; dc:title "On the Tityus forcipula species group: redescription of Tityus forcipula (Scorpiones, Buthidae), description of a new Andean species, and notes on the taxonomy of the group" ; fabio:hasPart , , , , , ; a fabio:JournalArticle . dwc:authority "(Gervais, 1843)" ; dwc:baseAuthorityName "Gervais" ; dwc:baseAuthorityYear "1843" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Gervais, 1843)" ; dwc:species "forcipula" ; dwc:subGenus "Atreus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Lourenco" ; dwc:authorityYear "2013" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Lourenco, 2013" ; dwc:species "crassicauda" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Lourenco" ; dwc:authorityYear "1994" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Lourenco, 1994" ; dwc:species "cuellari" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Kraepelin" ; dwc:baseAuthorityYear "1914" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Kraepelin, 1914)" ; dwc:species "fuhrmanni" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authority "Pinto-Da-Rocha & Cabra-García, 2022" ; dwc:authorityName "Pinto-Da-Rocha & Cabra-García" ; dwc:authorityYear "2022" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Pinto-Da-Rocha & Cabra-García, 2022" ; dwc:species "moralensis" ; dwc:subGenus "Atreus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Mello-Leitao" ; dwc:authorityYear "1945" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Mello-Leitao, 1945" ; dwc:species "dasyurus" ; dwc:subSpecies "fulvipes" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Borelli" ; dwc:authorityYear "1899" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Borelli, 1899" ; dwc:species "festae" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Lourenco" ; dwc:authorityYear "2000" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Lourenco, 2000" ; dwc:species "florezi" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Miranda, Bermudez, Florez D & de Armas" ; dwc:authorityYear "2020" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Miranda, Bermudez, Florez D & de Armas, 2020" ; dwc:species "jaimei" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Pocock" ; dwc:authorityYear "1897" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Pocock, 1897" ; dwc:species "macrochirus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Pocock" ; dwc:authorityYear "1897" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Pocock, 1897" ; dwc:species "metuendus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Pocock" ; dwc:authorityYear "1897" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Pocock, 1897" ; dwc:species "pachyurus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Gervais" ; dwc:baseAuthorityYear "1843" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Gervais, 1843)" ; dwc:species "obscurus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Pocock" ; dwc:authorityYear "1897" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:scientificNameAuthorship "Pocock, 1897" ; dwc:species "dasyurus" ; dwc:subSpecies "dasyurus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Perty" ; dwc:baseAuthorityYear "1833" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Perty, 1833)" ; dwc:species "bahiensis" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Kraepelin" ; dwc:authorityYear "1898" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Kraepelin, 1898" ; dwc:species "trivittatus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Thorell" ; dwc:baseAuthorityYear "1876" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Thorell, 1876)" ; dwc:species "stigmurus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:baseAuthorityName "Karsch" ; dwc:baseAuthorityYear "1879" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "(Karsch, 1879)" ; dwc:species "androcottoides" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Kraepelin" ; dwc:authorityYear "1895" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Kraepelin, 1895" ; dwc:species "bolivianus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "C. L. Koch" ; dwc:authorityYear "1844" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Koch, 1844" ; dwc:species "clathratus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Pocock" ; dwc:authorityYear "1893" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Pocock, 1893" ; dwc:species "asthenes" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:authorityName "Pocock" ; dwc:authorityYear "1897" ; dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Pocock, 1897" ; dwc:species "dasyurus" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "forcipula" ; dwc:subGenus "Atreus" ; trt:hasParentName , ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "crassicauda" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "cuellari" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "fuhrmanni" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "moralensis" ; dwc:subGenus "Atreus" ; trt:hasParentName , ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:species "dasyurus" ; dwc:subSpecies "fulvipes" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "dasyurus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "festae" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "florezi" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "jaimei" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "macrochirus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "metuendus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "pachyurus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "obscurus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "subSpecies" ; dwc:species "dasyurus" ; dwc:subSpecies "dasyurus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "bahiensis" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "trivittatus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "stigmurus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "androcottoides" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "bolivianus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "clathratus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "asthenes" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Arachnida" ; dwc:family "Buthidae" ; dwc:genus "Tityus" ; dwc:kingdom "Animalia" ; dwc:order "Scorpiones" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:subGenus "Atreus" ; trt:hasParentName ; a dwcFP:TaxonName . dc:description "FIGURE 2. Tityus (Atreus) forcipula (Gervais, 1843), habitus. A, B. Male (MUSENUV-Ar 438). A. Dorsal view. B. Ventral view. C, D. Female (MUSENUV-Ar 436). C. Dorsal view. D. Ventral view. Scale bar= 10 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 3. Tityus (Atreus) moralensis sp. nov., habitus. A, B. Male holotype (MUSENUV-Ar 2104). A. Dorsal view. B. Ventral view. C, D. Female paratype (ICN-1402). C. Dorsal view. D. Ventral view. Scale bar= 15 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 9. Tityus (Atreus) Gervais, 1843, pectinal area. A, B. Tityus (Atreus) forcipula (Gervais, 1843).A. Male (MUSENUV- Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 12. Tityus (Atreus) Gervais, 1843, metasoma, dorsal view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 13. Tityus (Atreus) Gervais, 1843, metasoma, ventral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436). C, D. Tityus (Atreus) moralensis sp. nov. C. Male holotype (MUSENUV-Ar 2104). D. Female paratype (ICN-1402). Scale bars= 5 mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 15. Tityus (Atreus) Gervais, 1843, telson, lateral view. A, B. Tityus (Atreus) forcipula (Gervais, 1843). A. Male (MUSENUV-Ar 438). B. Female (MUSENUV-Ar 436).C, D. Tityus(Atreus) moralensis sp. nov. C. Male holotype(MUSENUV- Ar 2104). D. Female paratype (ICN-1402). Scale bars= 1.5 mm." ; fabio:hasRepresentation ; a fabio:Figure .