Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species
FIGURE 2. Some diagnostic characters of the Lepanus ustulatus species group. A, Dorsal view of male protibia of Lepanus ustulatus; B, dorsal view of female protibia of Lepanus ustulatus; C, dorsal view of male protibia of Lepanus globulus; D, male metatibia of Lepanus ustulatus; E, female metatibia of Lepanus ustulatus; F, male metatibia of Lepanus lemannae; G, lateral view of male protibia of Lepanus cameroni; H, lateral view of male protibia of Lepanus globulus; I, lateral view of male protibia of Lepanus cardwellensis; J, head of Lepanus cameroni showing eye canthus not dividing the eye; K, posterolateral view of pronotum of Lepanus globulus showing corners distinct; L, head of Lepanus globulus showing eye canthus almost dividing the eye; M, posterolateral view of pronotum of Lepanus cardwellensis showing corners rounded.
FIGURE 7. A, Map of known records of Lepanus cardwellensis, Lepanus globulus, and Lepanus lemannae with the zones of insect endemism of the Wet Tropics sensu Yeates & Montieth (2008) numbered and indicated by dashed lines. Zone numbers are as follows: 1 Mount Finnigan; 2 Thornton Peak; 3 Windsor Tableland; 4 Carbine Tableland; 5 Hann Tableland; 6 Black Mountain; 7 Lamb Range; 8 Walsh/Hugh Nelson Range; 9 Atherton Tableland; 10 Bellenden Ker; 11 Malbon Thompson Range; 12 Walter Hill Range; 13 Kirrama/Cardwell Ranges; 14 Seaview Range; 15 Hinchinbrook Island; 16 Paluma/Bluewater Ranges; 17 Mount Elliot, B, predicted distribution of Lepanus globulus; C, predicted distribution of Lepanus lemannae.
FIGURE 1. Phylogenetic estimate of relationships among described species in the genus Lepanus. Consensus topology generated from Bayesian analysis (BA) with node values for posterior probability BA and bootstrap values of clades also supported in maximum-likelihood analysis. Strongly supported nodes are indicated by coloured circles at base of nodes, while other support values are listed in full. Species groups revised here are highlighted by the grey boxes and IBRA regions for these specimens are also listed following the general locality in bold. Abbreviations for localities are as follows CMC: Central Mackay Coast; CYP: Cape York Peninsula; NNC: NSW North Coast; NSW: New South Wales; NT: Northern Territory; QLD: Queensland; SEQ: South Eastern Queensland; SYB: Sydney Basin; TS: Torres Strait; WA: Western Australia; WET: Wet Tropics.