@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . cito:cites , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; dc:creator "Vilela, Diogo Silva; Lencioni, Frederico A. A.; Bota-Sierra, Cornelio A.; Ware, Jessica L.; Bispo, Pitágoras C." ; dc:title "Heteragrion , Selys 1862" ; trt:augmentsTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "96" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:pubDate "2023-10-16" ; bibo:startPage "1" ; bibo:volume "5356" ; dc:creator "Vilela, Diogo Silva; Lencioni, Frederico A. A.; Bota-Sierra, Cornelio A.; Ware, Jessica L.; Bispo, Pitágoras C." ; dc:date "2023" ; dc:title "Taxonomic revision of the Neotropical genus Heteragrion Selys, 1862 (Zygoptera: Heteragrionidae): male morphology, new species and illustrated key" ; fabio:hasPart , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , , ; a fabio:JournalArticle . dwc:authority "Selys, 1862" ; dwc:authorityName ", Selys" ; dwc:authorityYear "1862" ; dwc:class "Insecta" ; dwc:family "Megapodagrionidae" ; dwc:genus "Heteragrion" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Selys, 1862" ; rdfs:seeAlso ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Megapodagrionidae" ; dwc:genus "Heteragrion" ; dwc:kingdom "Animalia" ; dwc:order "Odonata" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Megapodagrionidae" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:order "Odonata" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "FIGURE 12. Heteragrion aurantiacum: (a–c) anterior view of head; (d–e) lateral view of pterothorax; (f–g, i) mediodorsal view of cercus; (h, j) close up on the apical portion of medial lobe (ML) showing some morphological variation." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 18. Heteragrion brianmayi: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 21. Heteragrion cauei: (a) anterior view of head; (b) lateral view of pterothorax; (c) dorsal view of cercus; (d–e) mediodorsal view of cercus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 11. Heteragrion archon: (a) anterior view of head; (b) lateral view of pterothorax; (c) lateral view of anal appendages; (d) mediodorsal view of anal appendages; (a–b) adapted from the original drawings by De Marmels (2008). Figures c–d provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 53. Heteragrion palmichale: (a) anterior view of head; (b) lateral view of pterothorax; (c) lateral view of anal appendages; (a–b) adapted from Hartung (2002)." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 1. The Heteragrion cercus, showing the three main regions of the appendage: basal portion (BP), medial portion (MP) and apical portion (AP). Note that the proportions between cercus (a) and (b) are quite different and serve as a useful tool for species identification. ML: medial lobe. Figure b provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 2. The Heteragrion cercus, showing some differences between the two groups of species: (a–d) Group A species, (e–h) Group B species. BP: cercus basal portion; ML: medial lobe. Figures a, g–h provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 3. The Heteragrion cercus, showing some differences between the two groups of species: (a–c) Group A species, (d–f) Group B species. ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 10. Heteragrion angustipenne: (a, c) anterior view of male head; (b) female head; (d, f) dorsal view of cercus; (e, g) mediodorsal view of cercus; (c) adapted from Williamson (1919), based on a male examined by Calvert (1909). ML: medial lobe. Figures f–g provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 14. Heteragrion bariai: (a–b) anterior view of head; (c) lateral view of pterothorax; (d, g) mediodorsal view of cercus; (e) medial view of cercus; (f) dorsal view of cercus. BP: basal portion; MP: medial portion; AP: apical portion; ML: medial lobe. Figures f–g provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 15. Heteragrion beschkii: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) lateral view of cercus. BP: basal portion." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 25. Heteragrion consors: (a) anterior view of head; (b) lateral view of pterothorax; (c–e) mediodorsal view of cercus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 52. Heteragrion ovatum: (a) lateral view of pterothorax; (b) anterior view of head, lateral view of thorax, and S1–2; (c–d) dorsal view of pterothorax; (e–f) mediodorsal view of cercus; (g–h) lateral view of cercus. BP: basal portion; MP: medial portion. Figures a–j provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 4. The Heteragrion cercus, showing some differences between the two groups of species: (a–c) Group A species, (d–f) Group B species. BP: basal portion. Figure d provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 51. Heteragrion muryense: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus. BP: basal portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 26. Heteragrion cooki: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus. ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 63. Heteragrion thais: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus; (a) adapted from Machado (2015). ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 65. Heteragrion triangulare: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 28. Heteragrion cyane: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) dorsal view of cercus. ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 45. Heteragrion lencionii: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus. ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 30. Heteragrion denisye: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) laterodorsal view of cerci. ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 37. Heteragrion gracile: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) close up of MP, showing the two-ridged ML. MP: medial portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 9. Heteragrion alienum: (a) anterior view of head; (b) lateral view of pterothorax; (c) dorsal view of cercus, showing longitudinal depression on AP; (d–e) cercus in mediodorsal view. AP: apical portion." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 20. Heteragrion calendulum: (a–c) anterior view of head; (d–e) mediodorsal view of cercus. MP: medial portion; (c) adapted from Stand-Pérez et al. (2019). AP: apical portion." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 32. Heteragrion erythrogastrum: (a) anterior view of head; (b) lateral view of pterothorax; (c) dorsal view of cercus; (d) mediodorsal view of cercus. ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 62. Heteragrion tatama: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of anal appendages; (e) dorsal view of anal appendages. MP: medial portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 19. Heteragrion calafatiensis: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) lateral view of cercus. AP: apical portion of cercus; MP: medial portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 8. Heteragrion albifrons: (a) anterior view of head; (b) lateral view of pterothorax; (c–e) cercus in mediodorsal view. BP: basal portion; MP: medial portion; AP: apical portion." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 31. Heteragrion eboratum: (a) anterior view of head; (b) dorsal view of anal appendages; (c) mediodorsal view of cercus; (a) adapted from the original drawings by Donnelly (1965). AP: apical portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 16. Heteragrion bickorum: (a–b) anterior view of head; (c) lateral view of pterothorax; (d) mediodorsal view of cercus; (e) close up of MP-AP in medial view, showing the elevated crest and the ridge above ML; (f) medial view of cercus; (g) dorsal view of cercus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 5. Some individual variation in coloration and morphology in Heteragrion species: H. ictericum, left column; H. silvarum, right column. Images provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 42. Heteragrion inca: (a–c) anterior view of head; (d) lateral view of pterothorax; (e–f, h) mediodorsal view of cercus; (g, i) close up showing the two MP ridges above ML; (c) adapted from the drawing by Williamson (1919). MP: medial portion of cercus; ML: medial lobe. Figures h–i provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 60. Heteragrion silvarum: (a–b) anterior view of head; (c) lateral view of pterothorax; (d–g) mediodorsal view of cercus. BP: basal portion; MP: medial portion; AP: apical portion." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 6. Map showing the distribution of Heteragrion species from Group A (green dots) and Group B (red dots). We used no more than five dots per species, in order not to cover the dots where the two groups overlapped in distribution." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 7. Heteragrion aequatoriale: (a–d) anterior view of head, showing minimal variation in color pattern; (e) cercus in dorsal view, showing the proportions of basal portion (BP), medial portion (MP), and apical portion (AP); (f) cercus in mediodorsal view; (g–h) close up of MP, showing ridge above medial lobe (ML) and small cleft at junction of ridge-ML. Figures e–h provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 55. Heteragrion peregrinum: (a–b) anterior view of head; (c–d) dorsal view of cerci; (e) mediodorsal view of cercus. BP: basal portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 38. Heteragrion ictericum: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) lateral view of cercus. BP: basal portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 54. Heteragrion pemon: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 24. Heteragrion cinnamomeum: (a) anterior view of head; (b) lateral view of pterothorax; (c–e) mediodorsal view of cercus. BP: basal portion; ML: medial lobe. Figure e provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 36. Heteragrion gorbi: (a) anterior view of head; (b) lateral view of pterothorax; (c–e) mediodorsal view of cercus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 27. Heteragrion corderoi sp. nov. male holotype: (a) dorsal view of head; (b) lateral view of habitus; (c) lateral view of pterothorax; (d) dorsal view of pterothorax (e–f) mediodorsal view of cercus. BP: basal portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 46. Heteragrion luizfelipei: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus; (e–f) close up of MP, showing the hollow space formed by a carina on ML. MP: medial portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 48. Heteragrion makiritare: (a) anterior view of head; (b) lateral view of pterothorax; (c) dorsal view of cercus; (d) mediodorsal view of cercus; (a–b) adapted from De Marmels (2004). ML: medial lobe. Figures c–d provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 49. Heteragrion mantiqueirae: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) lateral view of cercus; (a) adapted from Machado (2006). MP: medial portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 56. Heteragrion petiense: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) closeup of MP, showing ML plates. MP: medial portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 43. Heteragrion itacolomii: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus; (e) close up of MP, showing the ML plates. MP: medial portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 44. Heteragrion johndeaconi: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) lateral view of cercus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 35. Heteragrion freddiemercuryi: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) close up of MP, showing the ML plates. MP: medial portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 64. Heteragrion tiradentense: (a–b) anterior view of head; (c) lateral view of pterothorax; (d–f) mediodorsal view of cercus." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 57. Heteragrion rogertaylori: (a–b) anterior view of head; (c–d) lateral view of pterothorax; (e–g) mediodorsal view of cercus. ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 34. Heteragrion flavovittatum: (a) anterior view of head; (b) lateral view of pterothorax; (c–e) mediodorsal view of cercus. Figure e provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 58. Heteragrion roquei: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) lateral view of cercus. ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 17. Heteragrion breweri: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d) lateral view of anal appendages. ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 39. Heteragrion icterops: (a) anterior view of head; (b–c) lateral view of pterothorax; (d, f) mediodorsal view of cercus; (e) dorsal view of anal appendages. ML: medial lobe. Figures e–f provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 59. Heteragrion rubrifulvum: (a) anterior view of head; (b) lateral view of pterothorax; (c) dorsal view of anal appendages; (d) mediodorsal view of anal appendages; (a–b) adapted from Donnelly (1992). AP: apical portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 67. Heteragrion valgum: (a–b) anterior view of head; (c) lateral view of pterothorax; (d) dorsal view of cercus; (e) mediodorsal view of cercus; (b–c) adapted from Donnelly (1992). BP: basal portion." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 61. Heteragrion simulatum: (a–b) anterior view of head; (c) lateral view of pterothorax; (d) mediodorsal view of cercus; (e) lateral view of cercus; (a) adapted from Williamson (1919), (b–c) adapted from De Marmels (1989). ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 13. Heteragrion azulum: (a) anterior view of head; (b) lateral view of pterothorax; (c) dorsal view of cercus; (d) close up, MP-AP; (e) mediodorsal view of cercus; (f) close up, ridge and ML. MP: medial portion; AP: apical portions; ML: medial lobe. Figures a–f provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 23. Heteragrion chrysops: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus; (e) closeup of cercus BP in medial view. BP: basal portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 47. Heteragrion majus: (a) anterior view of head; (b) lateral view of pterothorax; (c, e) mediodorsal view of cercus; (d) dorsal view of cercus. ML: medial lobe; AP: apical portion. Figure e provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 22. Heteragrion chlorotaeniatum: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus; (e) dorsal view of cercus. BP: basal portion; ML medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 33. Heteragrion flavidorsum: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) medial view of cercus; (e–f) dorsal view of cercus; (g) mediodorsal view of cercus. BP: basal portion; MP: medial portion. Figures f–g provided by Rosser Garrison and Natalia von Ellenrieder." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 66. Heteragrion tricellulare: (a) anterior view of head; (b) lateral view of pterothorax; (c) mediodorsal view of cercus; (d–e) dorsal view of cercus. ML: medial lobe; AP: apical portion." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 29. Heteragrion demarmelsi: (a) anterior view of male head; (b) anterior view of female head; (c) mediodorsal view of cercus; (d) dorsal view of cercus; (e) lateral view of cercus; BP: basal portion; MP: medial portion; ML: medial lobe." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 50. Heteragrion mitratum: (a) anterior view of head; (b) lateral view of pterothorax; (c–d) mediodorsal view of cercus; (e) dorsal view of cercus. BP: basal portion; MP: medial portion; ML: medial lobe; AP: apical portion." ; fabio:hasRepresentation ; a fabio:Figure .