@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . @prefix xlink: . cito:cites , , , , , ; dc:creator "Shi, Hongliang; Liang, Hongbin" ; dc:title "Parena (Bothynoptera) heteronycha Shi & Liang 2023, sp. nov." ; dwc:basisOfRecord , , ; trt:definesTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "144" ; bibo:issue "1" ; bibo:journal "Zootaxa" ; bibo:pubDate "2023-05-18" ; bibo:startPage "1" ; bibo:volume "5286" ; dc:creator "Shi, Hongliang; Liang, Hongbin" ; dc:date "2023" ; dc:title "Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" ; fabio:hasPart , , , , , ; a fabio:JournalArticle . dwc:authority "Shi & Liang, 2023" ; dwc:authorityName "Shi & Liang" ; dwc:authorityYear "2023" ; dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:genus "Parena" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:scientificNameAuthorship "Shi & Liang, 2023" ; dwc:species "heteronycha" ; dwc:subGenus "Bothynoptera" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:genus "Parena" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "species" ; dwc:species "heteronycha" ; dwc:subGenus "Bothynoptera" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:genus "Parena" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "subGenus" ; dwc:subGenus "Bothynoptera" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:genus "Parena" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:family "Carabidae" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:order "Coleoptera" ; dwc:phylum "Arthropoda" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Insecta" ; dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:phylum "Arthropoda" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "FIGURE 26. Parena (Bothynoptera) heteronycha sp. n. A Habitus of holotype. B Habitus of paratype. C Mouthparts of holotype. D Labels of holotype. E Metatarsus. F Mesotarsus. G Protarsus. Scale bars = 2 mm for A, B; 0.5mm for E, F, G." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 11. Gonocoxites (left side) of ovipositor for Parena spp., ventral side (dorsal side for fig. Q). A–J subgenus Bothynoptera: A P. (Bothynoptera) heteronycha sp.n., Laos, paratype. B P. (Bothynoptera) dorsigera (Schaum), Shaanxi. C P. (Bothynoptera) kurosai Habu, Guangdong. D P. (Bothynoptera) tesari (Jedlička), Taiwan. E P. (Bothynoptera) taiwana Hua, Taiwan. F P. (Bothynoptera) tripunctata (Bates), Shaanxi. G P. (Bothynoptera) malaisei (Andrewes), Taiwan. H P. (Bothynoptera) quadrisignata Mateu, Yunnan. I P. (Bothynoptera) emarginata sp. n., holotype. J P. (Bothynoptera) triguttata sp. n., Sichuan, paratype. K–X subgenus Parena: K P. (Parena) bicolor Motschulsky, Sulawesi. L P. (Parena) rubripicta Andrewes, Yunnan. M P. (Parena) fasciata (Chaudoir), Taiwan. N P. (Parena) nigrolineata (Chaudoir), Taiwan. O P. (Parena) picipes sp. n., Borneo, paratype. P P. (Parena) andrewesi Jedlička, Philippine. Q P. (Parena) politissima (Chaudoir), Vanuatu. R P. (Parena) latecincta (Bates), Guizhou. S P. (Parena) circumdata Shibata, Taiwan. T P. (Parena) pendleburyi Andrewes, Borneo. U P. (Parena) fulva sp. n., Zimbabwe, paratype. V P. (Parena) madagascariensis (Alluaud), Madagascar. W P. (Parena) ruficornis sp.n., CentralAfrica, paratype. X P. (Parena) africana (Alluaud), Uganda. Y–γ subgenus Crossoglossa: Y P. (Crossoglossa) cavipennis (Bates), Taiwan. Z P. (Crossoglossa) laesipennis (Bates), Yunnan. α P. (Crossoglossa) testacea (Chaudoir), Zhejiang. β P. (Crossoglossa) sulawesiensis Kirschenhofer, Sulawesi. γ P. (Crossoglossa) sciakyi sp. n., holotype. Scale bar = 0.2mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 4. Elytral base (left) of Parena spp. A P. (Parena) bicolor Motschulsky, Sumatra. B P. (Parena) africana (Alluaud), Cameroon. C P. (Crossoglossa) laesipennis (Bates), Taiwan. D P. (Bothynoptera) tripunctata (Bates), Beijing. E P. (Bothynoptera) dorsigera (Schaum), Guangxi. F P. (Bothynoptera) heteronycha sp. n., Laos, paratype. Scale bars = 0.5mm (a for fig. D, b for figs A B E F, c for fig. C). Abbreviations: bp elytral basal pore; sa the apex of scutellum; d1 the first discal setigerous pore." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 5. Elytral apex (left) of Parena spp. A P. (Parena) africana (Alluaud), Cameroon. B P. (Parena) ruficornis sp. n., Central Africa, paratype. C P. (Parena) valeriae Facchini, holotype. D P. (Parena) madagascariensis (Alluaud), Madagascar. E P. (Bothynoptera) tesari (Jedlička), Taiwan. F P. (Bothynoptera) heteronycha sp. n., Laos, paratype. G P. (Bothynoptera) kurosai Habu, Japan. H P. (Bothynoptera) dorsigera (Schaum), Yunnan. I P. (Bothynoptera) dorsigera (Schaum), Beijing. J P. (Bothynoptera) dorsigera (Schaum), Japan. Scale bars = 0.5mm" ; fabio:hasRepresentation ; a fabio:Figure . dc:description "FIGURE 6. A Hind wing, P. (Crossoglossa) cavipennis (Bates), Beijing, scale bar = 1mm. Abbreviations: oc oblongum cell; wc wedge cell. B–G Tarsals claws of right hindlegs for Metallicina spp., scale bar = 0.2mm: B Pachycallida rufoplagiata Jeannel, Madagascar, Sept Lacs, female. C Metallica viridipennis Chaudoir, Cameroon, Bois des Singes, male. D P. (Crossoglossa) sciakyi sp. n., holotype. E P. (Bothynoptera) kurosai Habu, Nepal. F P. (Bothynoptera) heteronycha sp. n., Laos, paratype. G P. (Parena) latecincta (Bates), Vietnam. H Protibia of P. (Parena) nigrolineata (Chaudoir), scale bar = 0.5 mm. I Mesotarsomeres of male of P. (Parena) nigrolineata (Chaudoir), adhesive hairs present on mesotarsomere 1, scale bar = 0.2mm. J Mesotarsomeres of male for P. (Parena) latecincta (Bates), adhesive hairs absent from mesotarsomere 1, scale bar = 0.2mm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "MAP 5. Distributions for species in the Parena (Bothynoptera) heteronycha group and Parena (Bothynoptera) taiwana group." ; fabio:hasRepresentation ; a fabio:Figure . dwc:locality "Mt. Phu Pane" ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "1550" ; dwc:verbatimLatitude "20.2" ; dwc:verbatimLongitude "103.98333" ; trt:gbifOccurrenceId "4115871474" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2012-05-18" ; dwc:locality "MT. Phu Pane" ; dwc:recordedBy "Collectors Lgt." ; dwc:typeStatus "holotype" ; dwc:verbatimElevation "1550" ; dwc:verbatimLatitude "20.12" ; dwc:verbatimLongitude "103.59" ; trt:gbifOccurrenceId "4115871389" ; trt:httpUri ; a dwc:MaterialCitation . dwc:eventDate "2010-06-25" ; dwc:locality "LAOS" ; dwc:typeStatus "paratype" ; trt:gbifOccurrenceId "4115871317" ; trt:httpUri ; a dwc:MaterialCitation .