Santos, Wildio Ikaro Da Graça; Dolibaina, Diego Rodrigo; Dias, Fernando Maia Silva; Mielke, Olaf Hermann Hendrik; Casagrande, Mirna Martins
A review of the South American metalmark genus Alesa Doubleday, 1847 (Lepidoptera: Riodinidae: Eurybiini) with the description of two new species
Cramer, 1777 W. Santos, Dolibaina, Dias, O. Mielke & Casagrande
Godart, [1824] W. Santos, Dolibaina, Dias, O. Mielke & Casagrande
Santos & Dolibaina & Dias & Mielke & Casagrande, 2023
Santos & Dolibaina & Dias & Mielke & Casagrande
Santos, Dolibaina, Dias, Mielke & Casagrande, 2023
FIGURES 30–39. Males of species of “amesis group”, in dorsal and ventral views (left to right). 30–31, A. amesis (DZ 35.219) Brazil, Acre, Mâncio Lima. 32–33, A. mariae sp. nov. Peru, Madre de Dios: Puerto Maldonado (holotype). 34–35, A. lipara Brazil, Acre: Santa Rosa do Purus (DZ 35.246). 36–37, A. hemiurga Brazil, Amazonas: Tefé (DZ 34.976). 38–39, A. thelydrias Brazil, Pará: rio Cupari (holotype, NHMUK). Scale bar: 1cm.
FIGURES 40–47. Females of species of “amesis group”, in dorsal and ventral views (left to right). 40–41. A. amesis Brazil, Amazonas: Novo Air„o (DZ 35.186). 42–45, A. lipara Brazil,Amazonas: Fonte Boa. 42–43 DZ 48.237, 44–45 (USMN). 46–47, Putative female of A. hemiurga Brazil, Maranh„o: Açailândia (DZ 35.226). Scale bar: 1cm.
FIGURES 48–55. Wings upper side pattern with the main diagnostic characters of males (top) and females (bottom) belonging to species of the “amesis group”. 48, 53, A. amesis (male DZ 48.287, female DZ 35.186). 49, A. mariae sp. nov. (holotype). 50, 54, A. lipara (male DZ 35.246, female DZ 48.237). 51, 55, A. hemiurga (male DZ 34.976, putative female DZ 35.226). 52, A. thelydrias (holotype).
FIGURES 56–60. Underside pattern with the main diagnostic characters of the hindwing underside of males belonging to the “amesis group”. 56, A. amesis (DZ 48.287). 57, A. mariae sp. nov. (holotype). 58, A. lipara (DZ 35.246). 59, A. hemiurga (DZ 34.976). 60, A. thelydrias (holotype).
FIGURES 67–70. Male genitalia of species of “amesis group”. 67, A. amesis Brazil, Pará: Paragominas (DZ 35.209). 68, A. mariae sp. nov. Brazil, Acre: Mâncio Lima (DZ 35.036). 69, A. lipara Brazil, Acre: Mâncio Lima (DZ 35.126). 70, A. hemiurga Brazil, Acre: Bujari (DZ 35.106). Left to right: dorsal view of tegumen and uncus, lateral and ventral views of genitalia, and dorsal view of valvae and transtilla. Scale bar: 1mm.
FIGURES 71–76. Female genitalia of Alesa species. 71, A. prema Brazil, Mato Grosso: Barra do Garça (DZ 35.254). 72, A. juliae sp. nov. Brazil, Mato Grosso: Rio Arinos (DZ 35.144). 73, A. telephae Brazil, Acre: Mâncio Lima (DZ 35.094). 74, A. amesis Brazil, Amazonas: Tefé (DZ 35.116). 75, A. lipara Brazil, Amazonas: Fonte Boa (DZ 48.237). 76, Putative female of A. hemiurga Brazil, Amazonas: Tefé (DZ 35.266). Up to bottom: lateral view of papilla analis and ventral view of sterigma and bursa copulatrix. Scale bar: 1mm.
FIGURES 77–81. Signum of species of Alesa. 77, A. prema Brazil, Mato Grosso: Barra do Garça (DZ 35.254). 78, A. telephae Brazil, Acre: Mâncio Lima (DZ 35.094). 79, A. amesis Brazil, amazonas: Tefé (DZ 35.116). 80, A. lipara Brazil, Amazonas: Fonte Boa (DZ 48.237). 81, Putative female of A. hemiurga Brazil, Amazonas: Tefé (DZ 35.266). Scale bars: 1mm.
FIGURES 86–91. Species of Alesa in situ, part II. 86–88, A. amesis, males (86–87) and female (88). 89–91, A. lipara. (89–90) two couples, note the shiny non metalic aspect of the male wings upper side (90), the same female of the latter couple, upsidedown (91). All images from Brazil, Mato Grosso, Alta Floresta, Cristalino Lodge. Figs 86–88, 90–91 by Sidnei Dantas https:// www.inaturalist.org/people/siddantas, Fig. 89 by Will Carter https://calydna.com/index.php/taxon/4375-alesa-lipara.
FIGURES 95. Geographic distribution of Alesa amesis and A. mariae sp. nov.
Mielke & Casagrande & Carneiro & Dias & Dolibaina
Mielke & Casagrande & Carneiro & Dias & Dolibaina & Siewert & Salik
O. Mielke & Casagrande & Dolibaina & Medeiros