@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .
@prefix xlink: <http://www.w3.org/1999/xlink/> .

<http://treatment.plazi.org/id/3E5A87FCFFAB9D124824F94EFB15B63D>
    cito:cites <http://taxon-concept.plazi.org/id/Animalia/Parabrachyodus_Forster-Cooper_1915>, <http://taxon-concept.plazi.org/id/Animalia/Telmatodon_Pilgrim_1907>, <http://dx.doi.org/10.5281/zenodo.7927071>, <http://dx.doi.org/10.5281/zenodo.7927073>, <http://dx.doi.org/10.5281/zenodo.7927063>, <http://dx.doi.org/10.5281/zenodo.7927075>, <http://dx.doi.org/10.5281/zenodo.7927077>, <http://dx.doi.org/10.5281/zenodo.7927079>, <http://dx.doi.org/10.5281/zenodo.7927081> ;
    dc:creator "Gernelle, Killian; Lihoreau, Fabrice; Boisserie, Jean-Renaud; Marivaux, Laurent; MéTais, Grégoire; Antoine, Pierre-Olivier" ;
    dc:title "Parabrachyodus hyopotamoides" ;
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    trt:augmentsTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Parabrachyodus_hyopotamoides_Lydekker_1883> ;
    trt:citesTaxonName <http://taxon-name.plazi.org/id/Animalia/Brachyodus_giganteus>, <http://taxon-name.plazi.org/id/Animalia/Brachyodus_africanus>, <http://taxon-name.plazi.org/id/Animalia/Brachyodus_pilgrimi>, <http://taxon-name.plazi.org/id/Animalia/Parabrachyodus_obtusus>, <http://taxon-name.plazi.org/id/Animalia/Brachyodus_gandoiensis>, <http://taxon-name.plazi.org/id/Animalia/Brachyodus_platydens>, <http://taxon-name.plazi.org/id/Animalia/Brachyodus_orientalis>, <http://taxon-name.plazi.org/id/Animalia/Brachyodus_indicus>, <http://taxon-name.plazi.org/id/Animalia/Gonotelma_major> ;
    trt:deprecates <http://taxon-concept.plazi.org/id/Animalia/Anthracotherium_hyopotamoides_Lydekker_1883>, <http://taxon-concept.plazi.org/id/Animalia/Hyopotamus_giganteus_Lydekker_1883>, <http://taxon-concept.plazi.org/id/Animalia/Telmatodon_bugtiensis_Pilgrim_1907>, <http://taxon-concept.plazi.org/id/Animalia/Brachyodus_manchharensis_Prasad_1964> ;
    trt:publishedIn <http://dx.doi.org/10.1093/zoolinnean/zlac111> ;
    a trt:Treatment .

<http://dx.doi.org/10.1093/zoolinnean/zlac111>
    bibo:endPage "309" ;
    bibo:issue "1" ;
    bibo:journal "Zoological Journal of the Linnean Society" ;
    bibo:pubDate "2023-05-01" ;
    bibo:startPage "278" ;
    bibo:volume "198" ;
    dc:creator "Gernelle, Killian; Lihoreau, Fabrice; Boisserie, Jean-Renaud; Marivaux, Laurent; MéTais, Grégoire; Antoine, Pierre-Olivier" ;
    dc:date "2023" ;
    dc:title "New material of Parabrachyodus hyopotamoides from Samane Nala, Bugti Hills (Pakistan) and the origin of Merycopotamini (Mammalia: Hippopotamoidea)" ;
    fabio:hasPart <http://dx.doi.org/10.5281/zenodo.7927071>, <http://dx.doi.org/10.5281/zenodo.7927073>, <http://dx.doi.org/10.5281/zenodo.7927063>, <http://dx.doi.org/10.5281/zenodo.7927075>, <http://dx.doi.org/10.5281/zenodo.7927077>, <http://dx.doi.org/10.5281/zenodo.7927079>, <http://dx.doi.org/10.5281/zenodo.7927081> ;
    a fabio:JournalArticle .

<http://taxon-concept.plazi.org/id/Animalia/Parabrachyodus_hyopotamoides_Lydekker_1883>
    dwc:authority "(LYDEKKER, 1883)" ;
    dwc:baseAuthorityName "Lydekker" ;
    dwc:baseAuthorityYear "1883" ;
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Parabrachyodus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Artiodactyla" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "(Lydekker, 1883)" ;
    dwc:species "hyopotamoides" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Parabrachyodus_hyopotamoides> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Anthracotherium_hyopotamoides_Lydekker_1883>
    dwc:authority "Lydekker" ;
    dwc:authorityName "Lydekker" ;
    dwc:authorityYear "1883" ;
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Anthracotherium" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Artiodactyla" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Lydekker, 1883" ;
    dwc:species "hyopotamoides" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Anthracotherium_hyopotamoides> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Hyopotamus_giganteus_Lydekker_1883>
    dwc:authority "Lydekker" ;
    dwc:authorityName "Lydekker" ;
    dwc:authorityYear "1883" ;
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Hyopotamus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Artiodactyla" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Lydekker, 1883" ;
    dwc:species "giganteus" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Hyopotamus_giganteus> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Telmatodon_bugtiensis_Pilgrim_1907>
    dwc:authority "Pilgrim" ;
    dwc:authorityName "Pilgrim" ;
    dwc:authorityYear "1907" ;
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Telmatodon" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Artiodactyla" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Pilgrim, 1907" ;
    dwc:species "bugtiensis" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Telmatodon_bugtiensis> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Parabrachyodus_Forster-Cooper_1915>
    dwc:authority "Forster-Cooper" ;
    dwc:authorityName "Forster-Cooper" ;
    dwc:authorityYear "1915" ;
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Parabrachyodus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Artiodactyla" ;
    dwc:phylum "Chordata" ;
    dwc:rank "genus" ;
    dwc:scientificNameAuthorship "Forster-Cooper, 1915" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Parabrachyodus> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Brachyodus_manchharensis_Prasad_1964>
    dwc:authority "Prasad" ;
    dwc:authorityName "Prasad" ;
    dwc:authorityYear "1964" ;
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Brachyodus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Artiodactyla" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:scientificNameAuthorship "Prasad, 1964" ;
    dwc:species "manchharensis" ;
    trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Brachyodus_manchharensis> ;
    a dwcFP:TaxonConcept .

<http://taxon-concept.plazi.org/id/Animalia/Telmatodon_Pilgrim_1907>
    dwc:authority "Pilgrim" ;
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    dwc:authorityYear "1907" ;
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    dwc:genus "Telmatodon" ;
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    dwc:phylum "Chordata" ;
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    a dwcFP:TaxonConcept .

<http://taxon-name.plazi.org/id/Animalia/Parabrachyodus_hyopotamoides>
    dwc:class "Mammalia" ;
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    dwc:genus "Parabrachyodus" ;
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    dwc:phylum "Chordata" ;
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<http://taxon-name.plazi.org/id/Animalia/Parabrachyodus>
    dwc:class "Mammalia" ;
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    dwc:rank "genus" ;
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    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Anthracotheriidae>
    dwc:class "Mammalia" ;
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    dwc:kingdom "Animalia" ;
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<http://taxon-name.plazi.org/id/Animalia/Artiodactyla>
    dwc:class "Mammalia" ;
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<http://taxon-name.plazi.org/id/Animalia/Mammalia>
    dwc:class "Mammalia" ;
    dwc:kingdom "Animalia" ;
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<http://taxon-name.plazi.org/id/Animalia/Chordata>
    dwc:kingdom "Animalia" ;
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    dwc:rank "phylum" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia>
    dwc:kingdom "Animalia" ;
    dwc:rank "kingdom" ;
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<http://taxon-name.plazi.org/id/Animalia/Anthracotherium_hyopotamoides>
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Anthracotherium" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Artiodactyla" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:species "hyopotamoides" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Anthracotherium> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Anthracotherium>
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Anthracotherium" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Artiodactyla" ;
    dwc:phylum "Chordata" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Anthracotheriidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Hyopotamus_giganteus>
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Hyopotamus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Artiodactyla" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:species "giganteus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Hyopotamus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Hyopotamus>
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Hyopotamus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Artiodactyla" ;
    dwc:phylum "Chordata" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Anthracotheriidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Brachyodus_giganteus>
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Brachyodus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Artiodactyla" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:species "giganteus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Brachyodus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Brachyodus>
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Brachyodus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Artiodactyla" ;
    dwc:phylum "Chordata" ;
    dwc:rank "genus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Anthracotheriidae> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Brachyodus_africanus>
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Brachyodus" ;
    dwc:kingdom "Animalia" ;
    dwc:order "Artiodactyla" ;
    dwc:phylum "Chordata" ;
    dwc:rank "species" ;
    dwc:species "africanus" ;
    trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Brachyodus> ;
    a dwcFP:TaxonName .

<http://taxon-name.plazi.org/id/Animalia/Telmatodon_bugtiensis>
    dwc:class "Mammalia" ;
    dwc:family "Anthracotheriidae" ;
    dwc:genus "Telmatodon" ;
    dwc:kingdom "Animalia" ;
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    dwc:class "Mammalia" ;
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    dwc:class "Mammalia" ;
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    dwc:phylum "Chordata" ;
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    dwc:class "Mammalia" ;
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<http://dx.doi.org/10.5281/zenodo.7927071>
    dc:description "Figure 4. Fragmentary cranium of Parabrachyodus hyopotamoides from Samane Nala 4 (UM-SAM4-001) in A, lateral view with an interpretative drawing. The grey full lines represent the sutures between bone units or the demarcation of teeth crowns, the grey dotted lines concern depressions on the skull surface. B, ventral view, displaying P1/-P2/ alveoli, and P3/- M3/. Scale bars equal 50 mm." ;
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<http://dx.doi.org/10.5281/zenodo.7927073>
    dc:description "Figure 5. Upper teeth of Parabrachyodus hyopotamoides. A, left upper incisor (I2/ or I3/) in lingual (left) and labial (right) view (UM-SAM4-024). B, P1/-P4/ from a left maxilla (M12034) in occlusal view. C, right P1/ (UM-SAM4-025) in occlusal view (mirror). D, left P3/ (M10598) in occlusal view. E, left maxilla with P4/-M1/ in occlusal view, with a drawing of P4/. F, left dP3/ (M12820) in occlusal view. Abbreviations derived from the nomenclature introduced in Figure 2: protoc., protocrista. G, dental variation of Parabrachyodus hyopotamoides figured on composite upper molars (N = 26 for M3/; N = 11 for M2/). Variable characters are coloured. Character frequency: black: 100%; red: (75–99%); yellow: (50–74%); green: (25–49%); blue: (0–24%). The presence/absence of the coloured character is evaluated. 1, mesostyle divided into two slender apices; 2, parastyle of the same size or smaller than mesostyle; 3, wing-like labial cingulum; 4, postectoprotocrista reaching the lingual edge of the tooth; 5, double curvature of the postmetacristule. The arrow indicates the mesiolingual direction. Teeth are not to scale. H, left maxilla with M1/-M3/ (M12718) in occlusal view. I, right maxilla with P4/-M3/ (M12824) in occlusal view (mirror). Comparisons with drawings of (J) left M3/ of Brachyodus onoideus in occlusal view, modified from Ducrocq & Lihoreau (2006); K, left M3/ of Telmatodon orientalis (M12040, holotype) in occlusal view; L, right M3/ of Gonotelma shahbazi (UM-SAM4-010) in occlusal view (mirror); M, left M3/ of SiƲameryx palaeindicus (UM-SAM4-005) in occlusal view. N, M3/ of Par. hyopotamoides (UM-SAM5-001) in distal view, showing the low value of the angle between the lingual slope of the metaconule and the socket. Scale bars all equal 10 mm." ;
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<http://dx.doi.org/10.5281/zenodo.7927063>
    dc:description "Figure 1. Drawings from Forster-Cooper (1915) of A, the first upper dental row (P4/-M3/, M12031) and B, lower molar (M/3, M12030) allocated to the anthracotheriid Parabrachyodus, from the Bugti Hills, Pakistan. Note the high degree of wear and the focus of the author on P4/’s morphology. Scale bars equal 10 mm." ;
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<http://dx.doi.org/10.5281/zenodo.7927075>
    dc:description "Figure 6. Mandibles and lower teeth of Parabrachyodus hyopotamoides. A, right mandible with P/1-M/1 and symphysis (M12736) in occlusal view; 2nd for., second mandibular foramen; acc.cusp., accessory cuspids. B, left mandible with I/2-M/3/ and symphysis (M12724) in sagittal view of the symphysis with interpretative drawing; m.w., maximal width; C, frontal view" ;
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<http://dx.doi.org/10.5281/zenodo.7927077>
    dc:description "Figure 7. Hemi-mandible fragments of Parabrachyodus hyopotamoides from A, Tobah (Bugti Hills), right alveoli of P/3, and P/4-M/3 (UM-TOB-001) in occlusal view, and B, C, Safed Nala (Zinda Pir Dome), right M/3 (UM-SFN-001) in occlusal (B) and labial (C) views. Scale bars equal 10 mm." ;
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<http://dx.doi.org/10.5281/zenodo.7927079>
    dc:description "Figure 8. Enamel microstructure of Parabrachyodus hyopotamoides. A, vertical section of enamel from EDJ (bottom) to OES (top). B, detail of the decussation of several transition prisms in vertical section. C, detail of the angulation between IPM crystallites and prisms in vertical section. D, detail of the inter-row sheets in vertical section. EDJ, enamel dentine junction; OES, outer enamel surface; IPM, interprismatic matrix." ;
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<http://dx.doi.org/10.5281/zenodo.7927081>
    dc:description "Figure 9. Phylogenetic results. Strict consensus tree of the 208 most parsimonious trees of 1558 steps each (CI = 0.195, RI = 0.625). Letters above the branches at nodes refer to the clades discussed in the phylogenetic analysis section of the main text. Values under the branches at nodes are Bremer indices (BI) when> 1." ;
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