@prefix rdf: . @prefix rdfs: . @prefix bibo: . @prefix cito: . @prefix dc: . @prefix dwc: . @prefix dwcFP: . @prefix fabio: . @prefix trt: . # Warning: Failed to output a material citation, could not create identifier cito:cites , , , , , , , , ; dc:creator "Voris, Jared T.; Therrien, François; Zelenitsky, Darla K.; Brown, Caleb M." ; dc:title "Thanatotheristes Voris & Therrien & Zelenitsky & Brown 2020" ; trt:definesTaxonConcept ; trt:publishedIn ; a trt:Treatment . bibo:endPage "15" ; bibo:issue "104388" ; bibo:journal "Cretaceous Research" ; bibo:pubDate "2020-01-23" ; bibo:startPage "1" ; bibo:volume "110" ; dc:creator "Voris, Jared T.; Therrien, François; Zelenitsky, Darla K.; Brown, Caleb M." ; dc:date "2020" ; dc:title "A new tyrannosaurine (Theropoda: Tyrannosauridae) from the Campanian Foremost Formation of Alberta, Canada, provides insight into the evolution and biogeography of tyrannosaurids" ; fabio:hasPart , , , , , , , , ; a fabio:JournalArticle . dwc:authority "Voris & Therrien & Zelenitsky & Brown, 2020" ; dwc:authorityName "Voris & Therrien & Zelenitsky & Brown" ; dwc:authorityYear "2020" ; dwc:class "Reptilia" ; dwc:family "Tyrannosauridae" ; dwc:genus "Thanatotheristes" ; dwc:kingdom "Animalia" ; dwc:order "Dinosauria" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; dwc:scientificNameAuthorship "Voris, Therrien, Zelenitsky & Brown, 2020" ; trt:hasTaxonName ; a dwcFP:TaxonConcept . dwc:class "Reptilia" ; dwc:family "Tyrannosauridae" ; dwc:genus "Thanatotheristes" ; dwc:kingdom "Animalia" ; dwc:order "Dinosauria" ; dwc:phylum "Chordata" ; dwc:rank "genus" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:family "Tyrannosauridae" ; dwc:rank "family" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:order "Dinosauria" ; dwc:rank "order" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:class "Reptilia" ; dwc:rank "class" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:phylum "Chordata" ; dwc:rank "phylum" ; trt:hasParentName ; a dwcFP:TaxonName . dwc:kingdom "Animalia" ; dwc:rank "kingdom" ; a dwcFP:TaxonName . dc:description "Fig. 2. Skull reconstruction of Thanatotheristes holotype TMP 2010.5.7. Known bones appear in white. Missing bone morphologies and proportions are based on the holotype of Daspletosaurus torosus (CMN 8506). Scale bar equals 10 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig. 3. Locality of Thanatotheristes degrootorum holotype (TMP 2010.5.7).A) Geographic location of the locality.The specimen was found 0.5 km upstream from the confluence of the Bow and Oldman rivers. B) Photograph of locality looking NNE. Specimen was found scattered over a lateral extent of 10 m on the shore of the river. The nearby cliff is composed of coal beds and interbedded sandstones and mudstones of the Taber Coal Zone of the Foremost Formation. C) Stratigraphic section of the exposures near the fossil locality. Bracket indicates likely stratigraphic provenance of specimen. Dotted line marks the extent of the exposures at the type locality; the Herronton Sandstone is exposed farther downstream." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig. 4. Right maxilla of Thanatotheristes holotype (TMP 2010.5.7) in lateral view with A) known fragments overlain on hypothesized reconstruction and B) subcutaneous surface magnified. The subcutaneous surface is strongly ornamented and possesses autapomorphic dorsoventral ridges (arrows). Extent of antorbital fossa indicated by black dash line. Large healed bite mark is indicated by white dash line. Abbreviations: aofo, antorbital fossa; mf, maxillary fenestra; pmf, promaxillary fenestra; pnf, posterior-most neurovascular foramen; scs, subcutaneous surface; sppm, subnarial process of premaxilla. Scale bar equals 5 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig. 5. Left maxilla of referred specimen of Thanatotheristes (TMP 2018.16.1) in lateral view. Note the presence of pronounced vertical ridges (arrows) on subcutaneous surface, an autapomorphy of the taxon. Abbreviations: dvr, dorsoventral ridges; sc-afos, subcutaneous-antorbital fossa boundary. Scale bar equals 5 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig. 6. Comparison of right jugal (lateral view) and jugal sinus in tyrannosaurids. A) Thanatotheristes holotype (TMP 2010.5.7). B) Daspletosaurus torosus (TMP 2001.36.1).Black line represents position of transverse CT slice in the top right.Note the elliptical shape and extensive sinus inflation in the suborbital region cross section of Thanatotheristes, constrasting with the tear-drop shaped and constricted sinus of Daspletosaurus. Abbreviation: jse, jugal sinus endocast. Scale bars equal 3 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig. 7. Right postorbital fragment of Thanatotheristes holotype (TMP 2010.5.7). A) Specimen in lateral view, demonstrating arced subcutaneous ridges, B) and medial view.Note prominent bracing ridge for jugal contact on the medial surface of the bone. Abbreviations: br, bracing ridge for jugal contact; djc, dorsal extent of jugal contact; scr, subcutaneous ridges; sr, smooth region along lateral surface posterior margin. Scale bar equals 2 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig. 8. Comparison of left frontalin dorsalviewin tyrannosaurids.A)Thanatotheristes TMP 2010.5.7. White dash line represents nondiagnostic postorbital fragment attached to the frontal. B) Subcutaneous surface of frontal of Daspletosaurus torosus TMP 2001.036.0001. Abbreviations: apc, anterior postorbital contact expansion; aesc, autapomorphic anterior extension of sagittal crest; ifs, interfrontal suture;ls, lacrimal socket; np, nasal process; os, orbital slot; PA, parietal; PF, prefrontal; pfl, prefrontolacrimal process; pfn, prefrontonasal process; po, postorbital fragment; sc, sagittal crest; stf, supratemporal fossa; str, supratemporal ridge. Scale bars equal 5 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig. 9. Articulated right quadrate and surangular of Thanatotheristes holotype in lateral view. Abbreviations: anc, angular contact surface; ls, lateral shelf; ngf, non-glenoid fossa; pp, pneumatic pocket; qu, quadrate; sf, surangular foramen. Scale bar equals 2 cm." ; fabio:hasRepresentation ; a fabio:Figure . dc:description "Fig. 10. Right dentary of Thanatotheristes holotype (TMP 2010.5.7). A) Specimen in lateral view. B) Specimen in dorsal view. Numbers correspond to dentary alveolus position. Comparison of dentary symphyseal region between C) Thanatotheristes and D) Daspletosaurus torosus (TMP 2001.36.1). Note the absence of bony projections on the symphyseal surface and a rugose knob ventral to Meckelian foramen in Thanatotheristes contrasting with Daspletosaurus. Abbreviations: apr, anteroposterior ridges; bp, bony projections; dc, dentary chin; mf, meckelian foramen; mg, meckelian groove; rk, rugose knob. Scale bar equals 5 cm." ; fabio:hasRepresentation ; a fabio:Figure .