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@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .
@prefix xlink: <http://www.w3.org/1999/xlink/> .
<http://treatment.plazi.org/id/01B6AA753C235D27BFF5921091C3AAB0>
cito:cites <http://dx.doi.org/10.3897/zookeys.1209.125897.figure16>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure17>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure19>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure20>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure21>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure25> ;
dc:creator "Harden, Curt W.; Caterino, Michael S." ;
dc:title "Serranillus monadnock Harden & Caterino, 2024, sp. nov." ;
dwc:basisOfRecord <http://tb.plazi.org/GgServer/dwcaRecords/01B6AA753C235D27BFF5921091C3AAB0.mc.A6C21007563A8953974AC6717F43EA41>, <http://tb.plazi.org/GgServer/dwcaRecords/01B6AA753C235D27BFF5921091C3AAB0.mc.13248069306C88C2FFF343D5377268DD>, <http://tb.plazi.org/GgServer/dwcaRecords/01B6AA753C235D27BFF5921091C3AAB0.mc.150AA4B8E2D1EF9A422E45D40E8C049D>, <http://tb.plazi.org/GgServer/dwcaRecords/01B6AA753C235D27BFF5921091C3AAB0.mc.36E8180AC7DD06CDAEB747B9B34263F1> ;
trt:definesTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Serranillus_monadnock_Harden_2024> ;
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a trt:Treatment .
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bibo:endPage "197" ;
bibo:journal "ZooKeys" ;
bibo:pubDate "2024-08-08" ;
bibo:startPage "69" ;
bibo:volume "1209" ;
dc:creator "Harden, Curt W.; Caterino, Michael S." ;
dc:date "2024" ;
dc:title "Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)" ;
fabio:hasPart <http://dx.doi.org/10.3897/zookeys.1209.125897.figure16>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure17>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure19>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure20>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure21>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure25> ;
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dwc:authorityName "Harden & Caterino" ;
dwc:authorityYear "2024" ;
dwc:class "Insecta" ;
dwc:family "Carabidae" ;
dwc:genus "Serranillus" ;
dwc:kingdom "Animalia" ;
dwc:order "Coleoptera" ;
dwc:phylum "Arthropoda" ;
dwc:rank "species" ;
dwc:scientificNameAuthorship "Harden & Caterino, 2024" ;
dwc:species "monadnock" ;
trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Serranillus_monadnock> ;
a dwcFP:TaxonConcept .
<http://taxon-name.plazi.org/id/Animalia/Serranillus_monadnock>
dwc:class "Insecta" ;
dwc:family "Carabidae" ;
dwc:genus "Serranillus" ;
dwc:kingdom "Animalia" ;
dwc:order "Coleoptera" ;
dwc:phylum "Arthropoda" ;
dwc:rank "species" ;
dwc:species "monadnock" ;
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dwc:class "Insecta" ;
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dwc:kingdom "Animalia" ;
dwc:order "Coleoptera" ;
dwc:phylum "Arthropoda" ;
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trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Carabidae> ;
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dwc:class "Insecta" ;
dwc:family "Carabidae" ;
dwc:kingdom "Animalia" ;
dwc:order "Coleoptera" ;
dwc:phylum "Arthropoda" ;
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dwc:order "Coleoptera" ;
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dwc:kingdom "Animalia" ;
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dwc:kingdom "Animalia" ;
dwc:rank "kingdom" ;
a dwcFP:TaxonName .
<http://dx.doi.org/10.3897/zookeys.1209.125897.figure16>
dc:description "Figure 16. Distribution map of Serranillus species that occur in South Carolina. The locality for Serranillus sp. “ South Carolina, Coon Branch ” is the same as the single South Carolina S. jeanneli occurrence, and is not shown. Data are from Harden (2024)." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110355> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1209.125897.figure17>
dc:description "Figure 17. Multiple aspects of median lobes of South Carolina Serranillus species, progressively rotated dorsally from right lateral aspect (upper left) to left lateroventral aspect (lower right) A Serranillus dunavani B Serranillus jeanneli C Serranillus monadnock, holotype (Kings Mountain, SC) D Serranillus monadnock, paratype (Little Mountain, SC). Photographs not to same scale." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110356> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1209.125897.figure19>
dc:description "Figure 19. Serranillus monadnock holotype A dorsal habitus B abdominal ventrites, ventral aspect (black arrow = posterior margin of medial area with stretched microsculpture) C left paramere D right paramere E median lobe. Scale bars: 1 mm (A); 0.1 mm (B, C, D, E)." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110358> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1209.125897.figure20>
dc:description "Figure 20. Morphological features of Serranillus A Serranillus sp. “ South Carolina, Coon Branch ” dorsal habitus B head of Serranillus monadnock, dorsal aspect (black arrows = ocular tubercles) C dorsal aspect of abdominal ventrites of Serranillus septentrionis (black arrows = lateral extensions of last abdominal tergite). Scale bars: 1 mm (A); 0.1 mm (C)." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110359> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1209.125897.figure21>
dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110360> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1209.125897.figure25>
dc:description "Figure 25. Anillini habitat in South Carolina A Waldrop Stone Falls, Pickens Co. (Anillinus mica, Anillinus micamicus, Anillinus murrayae, Serranillus dunavani) B Lower Whitewater River gorge, Oconee Co. (Anillinus cherokee, A. murrayae, Anillinus sp. “ South Carolina, Coon Branch ”, S. dunavani, Serranillus jeanneli, Serranillus sp. “ South Carolina, Coon Branch ” C Long Cane Creek, Abbeville Co. (Anillinus chandleri, Anillinus dentatus, Anillinus jancae, S. dunavani). The holotypes of A. mica and A. micamicus were collected under embedded rocks uphill of the wooden staircase in A. The holotype of A. jancae was collected in a pipe trap set on a hill just out of view in C." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110364> ;
a fabio:Figure .
<http://tb.plazi.org/GgServer/dwcaRecords/01B6AA753C235D27BFF5921091C3AAB0.mc.A6C21007563A8953974AC6717F43EA41>
dwc:catalogNumber "ENT 00327997" ;
dwc:collectionCode "NCSU" ;
dwc:eventDate "1989-10-28" ;
dwc:typeStatus "holotype" ;
trt:httpUri <https://treatment.plazi.org/id/01B6AA753C235D27BFF5921091C3AAB0#A6C21007563A8953974AC6717F43EA41> ;
a dwc:MaterialCitation .
<http://tb.plazi.org/GgServer/dwcaRecords/01B6AA753C235D27BFF5921091C3AAB0.mc.13248069306C88C2FFF343D5377268DD>
dwc:collectionCode "NCSU" ;
dwc:locality "USA" ;
dwc:typeStatus "paratype" ;
trt:httpUri <https://treatment.plazi.org/id/01B6AA753C235D27BFF5921091C3AAB0#13248069306C88C2FFF343D5377268DD> ;
a dwc:MaterialCitation .
<http://tb.plazi.org/GgServer/dwcaRecords/01B6AA753C235D27BFF5921091C3AAB0.mc.150AA4B8E2D1EF9A422E45D40E8C049D>
dwc:catalogNumber "ENT 00327998, ENT 00327999" ;
dwc:collectionCode "NCSU" ;
dwc:eventDate "2007-12-08" ;
dwc:locality "Little Mountain" ;
dwc:recordedBy "J. and S. Cornell & D Pine Stump Hole" ;
dwc:typeStatus "paratype" ;
dwc:verbatimLatitude "34.188" ;
dwc:verbatimLongitude "-81.408" ;
trt:httpUri <https://treatment.plazi.org/id/01B6AA753C235D27BFF5921091C3AAB0#150AA4B8E2D1EF9A422E45D40E8C049D> ;
a dwc:MaterialCitation .
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dwc:catalogNumber "CUAC 000169317" ;
dwc:collectionCode "CUAC" ;
dwc:eventDate "2021-10-02" ;
dwc:locality "Coon Branch Natural Area" ;
dwc:recordedBy "C. W. Harden & On" ;
dwc:verbatimLatitude "35.0251" ;
dwc:verbatimLongitude "-83.0053" ;
trt:httpUri <https://treatment.plazi.org/id/01B6AA753C235D27BFF5921091C3AAB0#36E8180AC7DD06CDAEB747B9B34263F1> ;
a dwc:MaterialCitation .

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@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
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cito:cites <http://dx.doi.org/10.3897/zookeys.1209.125897.figure21>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure24>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure29>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure30>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure32> ;
dc:creator "Harden, Curt W.; Caterino, Michael S." ;
dc:title "Anillinus castaneus Harden & Caterino, 2024, sp. nov." ;
dwc:basisOfRecord <http://tb.plazi.org/GgServer/dwcaRecords/0CF12D7F8EEA59F88DAE48EB37D16388.mc.C870D705A930DE3DE23AD8C1A98052BB>, <http://tb.plazi.org/GgServer/dwcaRecords/0CF12D7F8EEA59F88DAE48EB37D16388.mc.2DB6CBE8D1EF5A226A04305E0A44D9A1>, <http://tb.plazi.org/GgServer/dwcaRecords/0CF12D7F8EEA59F88DAE48EB37D16388.mc.309F1744CB0A40DBBC23C32B25A834A7>, <http://tb.plazi.org/GgServer/dwcaRecords/0CF12D7F8EEA59F88DAE48EB37D16388.mc.FC2C9AEDD1A2E9ABA860F661470CB2F9>, <http://tb.plazi.org/GgServer/dwcaRecords/0CF12D7F8EEA59F88DAE48EB37D16388.mc.1145C7C9E77E8164D571937DB05B1304>, <http://tb.plazi.org/GgServer/dwcaRecords/0CF12D7F8EEA59F88DAE48EB37D16388.mc.809F1451AF36C990D25ECC7D944EB8B4>, <http://tb.plazi.org/GgServer/dwcaRecords/0CF12D7F8EEA59F88DAE48EB37D16388.mc.243A10B8DAD2DAC1711962C91D3A2430>, <http://tb.plazi.org/GgServer/dwcaRecords/0CF12D7F8EEA59F88DAE48EB37D16388.mc.944FC8D1838C3E47E0FB60D12FCDC5B0>, <http://tb.plazi.org/GgServer/dwcaRecords/0CF12D7F8EEA59F88DAE48EB37D16388.mc.D25AEE1A1EBA684F95F79A4576BE4003>, <http://tb.plazi.org/GgServer/dwcaRecords/0CF12D7F8EEA59F88DAE48EB37D16388.mc.187B2AD413F5E032F9FFB9885D9F4F8A> ;
trt:definesTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Anillinus_castaneus_Harden_2024> ;
trt:publishedIn <http://dx.doi.org/10.3897/zookeys.1209.125897> ;
a trt:Treatment .
<http://dx.doi.org/10.3897/zookeys.1209.125897>
bibo:endPage "197" ;
bibo:journal "ZooKeys" ;
bibo:pubDate "2024-08-08" ;
bibo:startPage "69" ;
bibo:volume "1209" ;
dc:creator "Harden, Curt W.; Caterino, Michael S." ;
dc:date "2024" ;
dc:title "Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)" ;
fabio:hasPart <http://dx.doi.org/10.3897/zookeys.1209.125897.figure21>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure24>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure29>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure30>, <http://dx.doi.org/10.3897/zookeys.1209.125897.figure32> ;
a fabio:JournalArticle .
<http://taxon-concept.plazi.org/id/Animalia/Anillinus_castaneus_Harden_2024>
dwc:authorityName "Harden & Caterino" ;
dwc:authorityYear "2024" ;
dwc:class "Insecta" ;
dwc:family "Carabidae" ;
dwc:genus "Anillinus" ;
dwc:kingdom "Animalia" ;
dwc:order "Coleoptera" ;
dwc:phylum "Arthropoda" ;
dwc:rank "species" ;
dwc:scientificNameAuthorship "Harden & Caterino, 2024" ;
dwc:species "castaneus" ;
trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Anillinus_castaneus> ;
a dwcFP:TaxonConcept .
<http://taxon-name.plazi.org/id/Animalia/Anillinus_castaneus>
dwc:class "Insecta" ;
dwc:family "Carabidae" ;
dwc:genus "Anillinus" ;
dwc:kingdom "Animalia" ;
dwc:order "Coleoptera" ;
dwc:phylum "Arthropoda" ;
dwc:rank "species" ;
dwc:species "castaneus" ;
trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Anillinus> ;
a dwcFP:TaxonName .
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dwc:class "Insecta" ;
dwc:family "Carabidae" ;
dwc:genus "Anillinus" ;
dwc:kingdom "Animalia" ;
dwc:order "Coleoptera" ;
dwc:phylum "Arthropoda" ;
dwc:rank "genus" ;
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dwc:class "Insecta" ;
dwc:family "Carabidae" ;
dwc:kingdom "Animalia" ;
dwc:order "Coleoptera" ;
dwc:phylum "Arthropoda" ;
dwc:rank "family" ;
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a dwcFP:TaxonName .
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dwc:class "Insecta" ;
dwc:kingdom "Animalia" ;
dwc:order "Coleoptera" ;
dwc:phylum "Arthropoda" ;
dwc:rank "order" ;
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dwc:class "Insecta" ;
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dwc:rank "class" ;
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dwc:kingdom "Animalia" ;
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dwc:rank "phylum" ;
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dwc:kingdom "Animalia" ;
dwc:rank "kingdom" ;
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<http://dx.doi.org/10.3897/zookeys.1209.125897.figure21>
dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110360> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1209.125897.figure24>
dc:description "Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110363> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1209.125897.figure29>
dc:description "Figure 29. Distribution map of valentinei - group species in South Carolina and adjacent states. Data are from Harden (2024)." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110368> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1209.125897.figure30>
dc:description "Figure 30. Aedeagi of valentinei group Anillinus species A – C Anillinus castaneus sp. nov. D – F Anillinus cornelli G – I Anillinus simplex sp. nov. J – N Anillinus murrayae. Median lobes C, F, I in right dorsolateral aspect, median lobe K in right lateral aspect. Left A, D, G, J and right B, E, H, L – N parameres in left and right lateral aspects, respectively. Scale bar: 0.1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110369> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1209.125897.figure32>
dc:description "Figure 32. Dorsal habitus and median lobes of Anillinus species A Anillinus castaneus sp. nov. Holotype B Anillinus sp. “ South Carolina, Chestnut Ridge ” C Anillinus cornelli, abdomen removed for DNA extraction. D A. castaneus sp. nov. Holotype, dorsal aspect E A. sp. “ South Carolina, Chestnut Ridge ”, dorsal aspect F A. cornelli, dorsal aspect. Black arrows point to proximal curve of flagellum. Scale bars: 1 mm (A – C); 0.1 mm (D – F)." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110371> ;
a fabio:Figure .
<http://tb.plazi.org/GgServer/dwcaRecords/0CF12D7F8EEA59F88DAE48EB37D16388.mc.C870D705A930DE3DE23AD8C1A98052BB>
dwc:catalogNumber "CUAC 000080962" ;
dwc:collectionCode "USNM" ;
dwc:eventDate "2018-04-08" ;
dwc:locality "Euparal" ;
dwc:recordedBy "Greenville Co. Chestnut Ridge Heritage Pres." ;
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dc:creator "Harden, Curt W.; Caterino, Michael S." ;
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dc:title "Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)" ;
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dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
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dc:description "Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." ;
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dc:description "Figure 30. Aedeagi of valentinei group Anillinus species A – C Anillinus castaneus sp. nov. D – F Anillinus cornelli G – I Anillinus simplex sp. nov. J – N Anillinus murrayae. Median lobes C, F, I in right dorsolateral aspect, median lobe K in right lateral aspect. Left A, D, G, J and right B, E, H, L – N parameres in left and right lateral aspects, respectively. Scale bar: 0.1 mm." ;
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dc:description "Figure 34. Dorsal habitus of Anillinus species in the elongatus group and sinuaticollis group A Anillinus montrex B Anillinus arenicollis C Anillinus choestoea sp. nov. D Anillinus seneca sp. nov. E Anillinus mica sp. nov. F Anillinus micamicus sp. nov. Abdomens removed in B – F for DNA extraction. Scale bars: 1 mm." ;
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dc:description "Figure 35. Male genitalia of Anillinus species in the sinuaticollis group A – C Anillinus choestoea sp. nov. D – F Anillinus seneca sp. nov. G – I Anillinus mica sp. nov. J – L Anillinus micamicus sp. nov. Median lobe of aedeagus in right dorsolateral (C, I, L) or right lateral (F) aspect. Left parameres (A, D, G, J) and right parameres (B, E, H, K) in left and right lateral aspects, respectively. Scale bar: 0.1 mm." ;
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dc:description "Figure 36. Distribution map of sinuaticollis - group species of Anillinus in South Carolina. Data are from Harden (2024). Blue lines show modern and pre-inundation courses of the Tugaloo River and Seneca River and their major tributaries. The confluence of the two rivers becomes the Savannah River." ;
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dc:description "Figure 25. Anillini habitat in South Carolina A Waldrop Stone Falls, Pickens Co. (Anillinus mica, Anillinus micamicus, Anillinus murrayae, Serranillus dunavani) B Lower Whitewater River gorge, Oconee Co. (Anillinus cherokee, A. murrayae, Anillinus sp. “ South Carolina, Coon Branch ”, S. dunavani, Serranillus jeanneli, Serranillus sp. “ South Carolina, Coon Branch ” C Long Cane Creek, Abbeville Co. (Anillinus chandleri, Anillinus dentatus, Anillinus jancae, S. dunavani). The holotypes of A. mica and A. micamicus were collected under embedded rocks uphill of the wooden staircase in A. The holotype of A. jancae was collected in a pipe trap set on a hill just out of view in C." ;
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dc:creator "Harden, Curt W.; Caterino, Michael S." ;
dc:title "Anillinus micamicus Harden & Caterino, 2024, sp. nov." ;
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bibo:endPage "197" ;
bibo:journal "ZooKeys" ;
bibo:pubDate "2024-08-08" ;
bibo:startPage "69" ;
bibo:volume "1209" ;
dc:creator "Harden, Curt W.; Caterino, Michael S." ;
dc:date "2024" ;
dc:title "Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)" ;
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dc:description "Figure 12. SEM micrographs of male left metafemora of Anillinus species. Scale bar: 0.1 mm." ;
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dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
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dc:description "Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." ;
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dc:description "Figure 25. Anillini habitat in South Carolina A Waldrop Stone Falls, Pickens Co. (Anillinus mica, Anillinus micamicus, Anillinus murrayae, Serranillus dunavani) B Lower Whitewater River gorge, Oconee Co. (Anillinus cherokee, A. murrayae, Anillinus sp. “ South Carolina, Coon Branch ”, S. dunavani, Serranillus jeanneli, Serranillus sp. “ South Carolina, Coon Branch ” C Long Cane Creek, Abbeville Co. (Anillinus chandleri, Anillinus dentatus, Anillinus jancae, S. dunavani). The holotypes of A. mica and A. micamicus were collected under embedded rocks uphill of the wooden staircase in A. The holotype of A. jancae was collected in a pipe trap set on a hill just out of view in C." ;
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dc:description "Figure 34. Dorsal habitus of Anillinus species in the elongatus group and sinuaticollis group A Anillinus montrex B Anillinus arenicollis C Anillinus choestoea sp. nov. D Anillinus seneca sp. nov. E Anillinus mica sp. nov. F Anillinus micamicus sp. nov. Abdomens removed in B – F for DNA extraction. Scale bars: 1 mm." ;
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dc:description "Figure 35. Male genitalia of Anillinus species in the sinuaticollis group A – C Anillinus choestoea sp. nov. D – F Anillinus seneca sp. nov. G – I Anillinus mica sp. nov. J – L Anillinus micamicus sp. nov. Median lobe of aedeagus in right dorsolateral (C, I, L) or right lateral (F) aspect. Left parameres (A, D, G, J) and right parameres (B, E, H, K) in left and right lateral aspects, respectively. Scale bar: 0.1 mm." ;
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dc:description "Figure 36. Distribution map of sinuaticollis - group species of Anillinus in South Carolina. Data are from Harden (2024). Blue lines show modern and pre-inundation courses of the Tugaloo River and Seneca River and their major tributaries. The confluence of the two rivers becomes the Savannah River." ;
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dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
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dc:description "Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." ;
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dc:description "Figure 25. Anillini habitat in South Carolina A Waldrop Stone Falls, Pickens Co. (Anillinus mica, Anillinus micamicus, Anillinus murrayae, Serranillus dunavani) B Lower Whitewater River gorge, Oconee Co. (Anillinus cherokee, A. murrayae, Anillinus sp. “ South Carolina, Coon Branch ”, S. dunavani, Serranillus jeanneli, Serranillus sp. “ South Carolina, Coon Branch ” C Long Cane Creek, Abbeville Co. (Anillinus chandleri, Anillinus dentatus, Anillinus jancae, S. dunavani). The holotypes of A. mica and A. micamicus were collected under embedded rocks uphill of the wooden staircase in A. The holotype of A. jancae was collected in a pipe trap set on a hill just out of view in C." ;
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dc:description "Figure 38. Distribution map of langdoni - group and loweae - group species that occur in South Carolina. Data are from Harden (2024)." ;
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dc:description "Figure 39. Dorsal habitus of female Anillinus in the loweae group A Anillinus merritti, sloped-humeri form (Georgia, Rabun Bald) B Anillinus loweae (Georgia, Rabun Cliffs) C Anillinus cherokee (South Carolina, Coon Branch). Scale bars: 1 mm." ;
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dwc:collectionCode "USNM" ;
dwc:eventDate "2004-04-05" ;
dwc:locality "Graham" ;
dwc:recordedBy "Berlese C. Carlton" ;
dwc:typeStatus "holotype" ;
dwc:verbatimLatitude "35.355553" ;
dwc:verbatimLongitude "-83.934166" ;
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dwc:catalogNumber "CUAC 000169308" ;
dwc:collectionCode "CUAC" ;
dwc:eventDate "2019-09-29" ;
dwc:locality "Rabun Bald" ;
dwc:recordedBy "C. W. Harden" ;
dwc:verbatimLatitude "34.9724" ;
dwc:verbatimLongitude "-83.302" ;
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dwc:catalogNumber "CUAC 000169307" ;
dwc:collectionCode "CUAC" ;
dwc:eventDate "2019-09-29" ;
dwc:locality "Rabun Bald" ;
dwc:recordedBy "C. W. Harden" ;
dwc:verbatimLatitude "34.9716" ;
dwc:verbatimLongitude "-83.3013" ;
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dwc:catalogNumber "CUAC 000169306" ;
dwc:collectionCode "CUAC" ;
dwc:eventDate "2019-10-26" ;
dwc:locality "Rabun Bald" ;
dwc:recordedBy "C. W. Harden" ;
dwc:verbatimLatitude "34.9748" ;
dwc:verbatimLongitude "-83.3059" ;
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dwc:collectionCode "CMNH" ;
dwc:eventDate "1964-05-30" ;
dwc:locality "Rabun Bald" ;
dwc:recordedBy "H. R. Steeves & J. D. Patrick & Jr." ;
dwc:verbatimLatitude "34.967" ;
dwc:verbatimLongitude "-83.299" ;
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dwc:collectionCode "TLc" ;
dwc:eventDate "2014-07-09" ;
dwc:locality "Rabun Bald" ;
dwc:recordedBy "T. Lawton" ;
dwc:verbatimLatitude "34.967" ;
dwc:verbatimLongitude "-83.299" ;
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dwc:catalogNumber "CUAC 000182308" ;
dwc:collectionCode "CUAC" ;
dwc:eventDate "2020-09-01" ;
dwc:locality "Chunky Gal" ;
dwc:recordedBy "P. Wooden & F. Etzler" ;
dwc:verbatimLatitude "35.1467" ;
dwc:verbatimLongitude "-83.7146" ;
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dwc:eventDate "2021-07-06" ;
dwc:locality "Chunky Gal" ;
dwc:recordedBy "M. S. Caterino" ;
dwc:verbatimLatitude "35.1471" ;
dwc:verbatimLongitude "-83.7144" ;
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dwc:eventDate "2019-07-19" ;
dwc:locality "Teyahalee Bald" ;
dwc:verbatimLatitude "35.2585" ;
dwc:verbatimLongitude "-83.7959" ;
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dwc:eventDate "2022-04-12" ;
dwc:locality "Teyahalee Bald" ;
dwc:verbatimLatitude "35.2598" ;
dwc:verbatimLongitude "-83.797" ;
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dwc:eventDate "2020-09-27" ;
dwc:locality "Stratton Ridge" ;
dwc:recordedBy "C. W. Harden" ;
dwc:verbatimLatitude "35.3382" ;
dwc:verbatimLongitude "-84.0249" ;
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dwc:catalogNumber "CUAC 000169309" ;
dwc:collectionCode "CUAC" ;
dwc:eventDate "2020-05-28" ;
dwc:locality "Stratton Ridge" ;
dwc:recordedBy "C. W. Harden" ;
dwc:verbatimLatitude "35.339" ;
dwc:verbatimLongitude "-84.025" ;
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dwc:collectionCode "CMNH" ;
dwc:eventDate "1974-08-24" ;
dwc:locality "Tapoco" ;
dwc:recordedBy "J. L. Bengston" ;
dwc:verbatimLatitude "35.357" ;
dwc:verbatimLongitude "-83.933" ;
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dwc:catalogNumber "ENT 00293740" ;
dwc:collectionCode "NCSU" ;
dwc:eventDate "2007-10-19" ;
dwc:locality "Wright Creek" ;
dwc:recordedBy "I. Sokolov" ;
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dwc:catalogNumber "CUAC 000170595" ;
dwc:collectionCode "CUAC" ;
dwc:eventDate "2015-06-29" ;
dwc:locality "Ellicott Rock" ;
dwc:recordedBy "S. Myers" ;
dwc:verbatimLatitude "35.0029" ;
dwc:verbatimLongitude "-83.1094" ;
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dwc:catalogNumber "CUAC 000169311, CUAC 000170594" ;
dwc:collectionCode "CUAC" ;
dwc:eventDate "2015-07-18" ;
dwc:locality "Ellicott Rock" ;
dwc:recordedBy "S. Myers" ;
dwc:verbatimLatitude "35.0075" ;
dwc:verbatimLongitude "-83.1358" ;
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dwc:collectionCode "CMNH" ;
dwc:eventDate "1965-08-08" ;
dwc:locality "Highlands" ;
dwc:recordedBy "H. R. Steeves" ;
dwc:verbatimLatitude "35.045" ;
dwc:verbatimLongitude "-83.451" ;
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dwc:eventDate "1965-05-22" ;
dwc:locality "Highlands" ;
dwc:recordedBy "H. R. Steeves" ;
dwc:verbatimLatitude "35.045" ;
dwc:verbatimLongitude "-83.451" ;
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dwc:eventDate "1965-05-23" ;
dwc:locality "Highlands" ;
dwc:recordedBy "H. R. Steeves" ;
dwc:verbatimLatitude "35.045" ;
dwc:verbatimLongitude "-83.451" ;
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dwc:locality "Highlands" ;
dwc:recordedBy "H. R. Steeves" ;
dwc:verbatimLatitude "35.045" ;
dwc:verbatimLongitude "-83.451" ;
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dwc:eventDate "1973-06-08" ;
dwc:locality "Coweeta Hydrological Lab" ;
dwc:recordedBy "W. Suter" ;
dwc:verbatimLatitude "35.045" ;
dwc:verbatimLongitude "-85.451" ;
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dwc:eventDate "1981-07-07" ;
dwc:locality "Highlands" ;
dwc:recordedBy "J. Pakaluk" ;
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dwc:eventDate "1970-08-08" ;
dwc:locality "Highlands" ;
dwc:recordedBy "T. C. Barr" ;
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dwc:eventDate "1949-06-27" ;
dwc:locality "Cullasaja River" ;
dwc:recordedBy "J. M. Valentine" ;
dwc:verbatimLatitude "35.067" ;
dwc:verbatimLongitude "-83.238" ;
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dwc:catalogNumber "CUAC 000169310" ;
dwc:collectionCode "CUAC" ;
dwc:eventDate "2015-07-22" ;
dwc:locality "Jones Gap" ;
dwc:recordedBy "S. Myers" ;
dwc:verbatimLatitude "35.0785" ;
dwc:verbatimLongitude "-83.2923" ;
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dwc:eventDate "1982-08-22" ;
dwc:locality "Buckhorn Gap" ;
dwc:recordedBy "J. Pakaluk" ;
dwc:verbatimLatitude "35.085" ;
dwc:verbatimLongitude "-83.26" ;
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dwc:eventDate "1971-05-14" ;
dwc:locality "Ridge" ;
dwc:recordedBy "T. C. Barr" ;
dwc:verbatimLatitude "35.11" ;
dwc:verbatimLongitude "-83.271" ;
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dwc:eventDate "1971-05-15" ;
dwc:locality "Old Road Gap" ;
dwc:recordedBy "T. C. Barr" ;
dwc:verbatimLatitude "35.173" ;
dwc:verbatimLongitude "-83.726" ;
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dwc:catalogNumber "CUAC 000182306, CUAC 000182307" ;
dwc:collectionCode "CUAC" ;
dwc:eventDate "2020-09-15" ;
dwc:locality "Copper Ridge Bald" ;
dwc:recordedBy "F. Etzler" ;
dwc:verbatimLatitude "35.2348" ;
dwc:verbatimLongitude "-83.5596" ;
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dwc:catalogNumber "CUAC 000182309" ;
dwc:collectionCode "CUAC" ;
dwc:eventDate "2020-09-15" ;
dwc:locality "Cowee Bald" ;
dwc:recordedBy "F. Etzler" ;
dwc:verbatimLatitude "35.3269" ;
dwc:verbatimLongitude "-83.335" ;
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dwc:catalogNumber "CUAC 000169305" ;
dwc:collectionCode "CUAC" ;
dwc:eventDate "2021-03-30" ;
dwc:locality "Doran Creek" ;
dwc:recordedBy "C. W. Harden & L. M. Thompson" ;
dwc:verbatimLatitude "34.7512" ;
dwc:verbatimLongitude "-83.2244" ;
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dwc:catalogNumber "LSAM 0295327, LSAM 0295330" ;
dwc:collectionCode "LSAM" ;
dwc:eventDate "1983-05-29" ;
dwc:locality "Highway" ;
dwc:recordedBy "D. S. Chandler" ;
dwc:verbatimLatitude "34.942" ;
dwc:verbatimLongitude "-83.089" ;
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dc:description "Figure 10. SEM micrographs of male protarsi A Anillinus jancae, left protarsus, dorsal aspect B Anillinus mica, left protarsus, dorsal aspect C Anillinus montrex left protarsus, dorsal aspect D A. jancae, right protarsus, ventral aspect E Anillinus choestoea, right protarsus, ventral aspect F Anillinus dentatus, right protarsus, ventral aspect G Serranillus dunavani (Jeannel), right protarsus, ventral aspect H Anillinus murrayae Sokolov & Carlton, right protarsus, ventral aspect I A. montrex, right protarsus, ventral aspect. Scale bars: 0.1 mm." ;
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dc:description "Figure 11. SEM micrographs of mouthparts of Appalachian Anillini genera A mentum of Serranillus dunavani (Jeannel) B mentum of Anillinus choestoea C mandibles of S. dunavani, dorsal aspect D mandibles of A. choestoea, dorsal aspect. Scale bars: 0.1 mm." ;
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dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
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dc:description "Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." ;
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dc:description "Figure 34. Dorsal habitus of Anillinus species in the elongatus group and sinuaticollis group A Anillinus montrex B Anillinus arenicollis C Anillinus choestoea sp. nov. D Anillinus seneca sp. nov. E Anillinus mica sp. nov. F Anillinus micamicus sp. nov. Abdomens removed in B – F for DNA extraction. Scale bars: 1 mm." ;
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dc:description "Figure 35. Male genitalia of Anillinus species in the sinuaticollis group A – C Anillinus choestoea sp. nov. D – F Anillinus seneca sp. nov. G – I Anillinus mica sp. nov. J – L Anillinus micamicus sp. nov. Median lobe of aedeagus in right dorsolateral (C, I, L) or right lateral (F) aspect. Left parameres (A, D, G, J) and right parameres (B, E, H, K) in left and right lateral aspects, respectively. Scale bar: 0.1 mm." ;
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dc:description "Figure 36. Distribution map of sinuaticollis - group species of Anillinus in South Carolina. Data are from Harden (2024). Blue lines show modern and pre-inundation courses of the Tugaloo River and Seneca River and their major tributaries. The confluence of the two rivers becomes the Savannah River." ;
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dc:description "Figure 15. Serranillus dunavani (Jeannel) A habitus B last abdominal ventrite, ventral aspect C left paramere, left lateral aspect D right paramere, right lateral aspect E median lobe, right dorsolateral aspect. Scale bars: 1 mm (A); 0.1 mm (B – E)." ;
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dc:description "Figure 18. Serranillus jeanneli Barr. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C left paramere, left lateral aspect D median lobe (with right paramere attached, black arrow), right dorsolateral aspect. Scale bars: 1 mm (A); 0.1 mm (B – D)." ;
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dc:description "Figure 19. Serranillus monadnock holotype A dorsal habitus B abdominal ventrites, ventral aspect (black arrow = posterior margin of medial area with stretched microsculpture) C left paramere D right paramere E median lobe. Scale bars: 1 mm (A); 0.1 mm (B, C, D, E)." ;
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dc:description "Figure 20. Morphological features of Serranillus A Serranillus sp. “ South Carolina, Coon Branch ” dorsal habitus B head of Serranillus monadnock, dorsal aspect (black arrows = ocular tubercles) C dorsal aspect of abdominal ventrites of Serranillus septentrionis (black arrows = lateral extensions of last abdominal tergite). Scale bars: 1 mm (A); 0.1 mm (C)." ;
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dc:description "Figure 11. SEM micrographs of mouthparts of Appalachian Anillini genera A mentum of Serranillus dunavani (Jeannel) B mentum of Anillinus choestoea C mandibles of S. dunavani, dorsal aspect D mandibles of A. choestoea, dorsal aspect. Scale bars: 0.1 mm." ;
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dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
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dc:description "Figure 14. Details of late-instar larvae of Appalachian Anillini A head of Anillinus jancae sp. nov., dorsal aspect, left antenna and right mandible omitted B left mandible of Serranillus dunavani (Jeannel), dorsal aspect C right maxilla of A. jancae, dorsal aspect D right maxilla of S. dunavani, dorsal aspect E urogomphi of A. jancae, dorsal aspect. Scale bar: 0.1 mm." ;
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dc:description "Figure 1. Distribution maps of eastern Nearctic Anillini A Serranillus B Anillinus. White dots: all known occurrences. Black dots: locations of new DNA sequence vouchers from this study. Data from Harden (2024)." ;
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dc:creator "Harden, Curt W.; Caterino, Michael S." ;
dc:date "2024" ;
dc:title "Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)" ;
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dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
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dc:description "Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." ;
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dc:description "Figure 38. Distribution map of langdoni - group and loweae - group species that occur in South Carolina. Data are from Harden (2024)." ;
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dc:description "Figure 39. Dorsal habitus of female Anillinus in the loweae group A Anillinus merritti, sloped-humeri form (Georgia, Rabun Bald) B Anillinus loweae (Georgia, Rabun Cliffs) C Anillinus cherokee (South Carolina, Coon Branch). Scale bars: 1 mm." ;
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dc:description "Figure 40. Anillinus sp. “ South Carolina, Wateree. ” A dorsal habitus B left metafemora, ventral aspect (black arrow = metafemoral spine) C spermatheca within cleared abdomen. Scale bars: 1 mm (A) and 0.1 mm (B, C)." ;
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dc:description "Figure 13. Dorsal habitus of late-instar larvae A Serranillus dunavani (Jeannel) B Anillinus jancae. Scale bars: 1 mm." ;
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dc:description "Figure 14. Details of late-instar larvae of Appalachian Anillini A head of Anillinus jancae sp. nov., dorsal aspect, left antenna and right mandible omitted B left mandible of Serranillus dunavani (Jeannel), dorsal aspect C right maxilla of A. jancae, dorsal aspect D right maxilla of S. dunavani, dorsal aspect E urogomphi of A. jancae, dorsal aspect. Scale bar: 0.1 mm." ;
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dc:description "Figure 30. Aedeagi of valentinei group Anillinus species A – C Anillinus castaneus sp. nov. D – F Anillinus cornelli G – I Anillinus simplex sp. nov. J – N Anillinus murrayae. Median lobes C, F, I in right dorsolateral aspect, median lobe K in right lateral aspect. Left A, D, G, J and right B, E, H, L – N parameres in left and right lateral aspects, respectively. Scale bar: 0.1 mm." ;
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dc:description "Figure 32. Dorsal habitus and median lobes of Anillinus species A Anillinus castaneus sp. nov. Holotype B Anillinus sp. “ South Carolina, Chestnut Ridge ” C Anillinus cornelli, abdomen removed for DNA extraction. D A. castaneus sp. nov. Holotype, dorsal aspect E A. sp. “ South Carolina, Chestnut Ridge ”, dorsal aspect F A. cornelli, dorsal aspect. Black arrows point to proximal curve of flagellum. Scale bars: 1 mm (A – C); 0.1 mm (D – F)." ;
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dc:title "Anillinus mica Harden & Caterino, 2024, sp. nov." ;
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bibo:pubDate "2024-08-08" ;
bibo:startPage "69" ;
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dc:creator "Harden, Curt W.; Caterino, Michael S." ;
dc:date "2024" ;
dc:title "Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)" ;
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dc:description "Figure 10. SEM micrographs of male protarsi A Anillinus jancae, left protarsus, dorsal aspect B Anillinus mica, left protarsus, dorsal aspect C Anillinus montrex left protarsus, dorsal aspect D A. jancae, right protarsus, ventral aspect E Anillinus choestoea, right protarsus, ventral aspect F Anillinus dentatus, right protarsus, ventral aspect G Serranillus dunavani (Jeannel), right protarsus, ventral aspect H Anillinus murrayae Sokolov & Carlton, right protarsus, ventral aspect I A. montrex, right protarsus, ventral aspect. Scale bars: 0.1 mm." ;
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dc:description "Figure 12. SEM micrographs of male left metafemora of Anillinus species. Scale bar: 0.1 mm." ;
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dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
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dc:description "Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." ;
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dc:description "Figure 25. Anillini habitat in South Carolina A Waldrop Stone Falls, Pickens Co. (Anillinus mica, Anillinus micamicus, Anillinus murrayae, Serranillus dunavani) B Lower Whitewater River gorge, Oconee Co. (Anillinus cherokee, A. murrayae, Anillinus sp. “ South Carolina, Coon Branch ”, S. dunavani, Serranillus jeanneli, Serranillus sp. “ South Carolina, Coon Branch ” C Long Cane Creek, Abbeville Co. (Anillinus chandleri, Anillinus dentatus, Anillinus jancae, S. dunavani). The holotypes of A. mica and A. micamicus were collected under embedded rocks uphill of the wooden staircase in A. The holotype of A. jancae was collected in a pipe trap set on a hill just out of view in C." ;
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dc:description "Figure 34. Dorsal habitus of Anillinus species in the elongatus group and sinuaticollis group A Anillinus montrex B Anillinus arenicollis C Anillinus choestoea sp. nov. D Anillinus seneca sp. nov. E Anillinus mica sp. nov. F Anillinus micamicus sp. nov. Abdomens removed in B – F for DNA extraction. Scale bars: 1 mm." ;
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dc:description "Figure 35. Male genitalia of Anillinus species in the sinuaticollis group A – C Anillinus choestoea sp. nov. D – F Anillinus seneca sp. nov. G – I Anillinus mica sp. nov. J – L Anillinus micamicus sp. nov. Median lobe of aedeagus in right dorsolateral (C, I, L) or right lateral (F) aspect. Left parameres (A, D, G, J) and right parameres (B, E, H, K) in left and right lateral aspects, respectively. Scale bar: 0.1 mm." ;
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dc:description "Figure 36. Distribution map of sinuaticollis - group species of Anillinus in South Carolina. Data are from Harden (2024). Blue lines show modern and pre-inundation courses of the Tugaloo River and Seneca River and their major tributaries. The confluence of the two rivers becomes the Savannah River." ;
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dc:title "Serranillus jeanneli Barr 1995" ;
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dc:creator "Harden, Curt W.; Caterino, Michael S." ;
dc:date "2024" ;
dc:title "Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)" ;
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dc:description "Figure 2. Habitus of eastern Nearctic Anillini. A Serranillus dunavani (South Carolina, Oconee Co.) B Anillinus moseleyae (Tennessee, Sevier Co.) C Anillinus dentatus (South Carolina, Abbeville Co.) D Serranillus jeanneli (South Carolina, Oconee Co.) E Anillinus folkertsioides (Alabama, Blount Co.) F Anillinus pecki (North Carolina, Avery Co.) G Anillinus sp. “ Georgia, Barnes Creek sp. 1 ” H Anillinus montrex (South Carolina, York Co.) I Anillinus sp. “ Tennessee, Kings Saltpeter Cave ” J Anillinus virginiae (Virginia, Bath Co.) K Anillinus erwini (Virginia, Grayson Co.) L Serranillus sp. (North Carolina, Graham Co.) M Anillinus sp. “ South Carolina, Waldrop Stone ” (South Carolina, Pickens Co.) N Anillinus sp. “ Alabama, Horseshoe Cave sp. 1 ” O Anillinus loweae (North Carolina, Haywood Co.) P Anillinus sp. “ Kentucky, Hestand sp. 1 ” Q Anillinus indianae (Indiana, Lawrence Co.) R Anillinus cf. barberi (West Virginia, Greenbrier Co.) S Anillinus hirsutus (Alabama, Madison Co.). Scale bar: 1 mm." ;
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dc:description "Figure 16. Distribution map of Serranillus species that occur in South Carolina. The locality for Serranillus sp. “ South Carolina, Coon Branch ” is the same as the single South Carolina S. jeanneli occurrence, and is not shown. Data are from Harden (2024)." ;
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dc:description "Figure 17. Multiple aspects of median lobes of South Carolina Serranillus species, progressively rotated dorsally from right lateral aspect (upper left) to left lateroventral aspect (lower right) A Serranillus dunavani B Serranillus jeanneli C Serranillus monadnock, holotype (Kings Mountain, SC) D Serranillus monadnock, paratype (Little Mountain, SC). Photographs not to same scale." ;
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dc:description "Figure 18. Serranillus jeanneli Barr. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C left paramere, left lateral aspect D median lobe (with right paramere attached, black arrow), right dorsolateral aspect. Scale bars: 1 mm (A); 0.1 mm (B – D)." ;
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dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
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dc:description "Figure 25. Anillini habitat in South Carolina A Waldrop Stone Falls, Pickens Co. (Anillinus mica, Anillinus micamicus, Anillinus murrayae, Serranillus dunavani) B Lower Whitewater River gorge, Oconee Co. (Anillinus cherokee, A. murrayae, Anillinus sp. “ South Carolina, Coon Branch ”, S. dunavani, Serranillus jeanneli, Serranillus sp. “ South Carolina, Coon Branch ” C Long Cane Creek, Abbeville Co. (Anillinus chandleri, Anillinus dentatus, Anillinus jancae, S. dunavani). The holotypes of A. mica and A. micamicus were collected under embedded rocks uphill of the wooden staircase in A. The holotype of A. jancae was collected in a pipe trap set on a hill just out of view in C." ;
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dc:description "Figure 2. Habitus of eastern Nearctic Anillini. A Serranillus dunavani (South Carolina, Oconee Co.) B Anillinus moseleyae (Tennessee, Sevier Co.) C Anillinus dentatus (South Carolina, Abbeville Co.) D Serranillus jeanneli (South Carolina, Oconee Co.) E Anillinus folkertsioides (Alabama, Blount Co.) F Anillinus pecki (North Carolina, Avery Co.) G Anillinus sp. “ Georgia, Barnes Creek sp. 1 ” H Anillinus montrex (South Carolina, York Co.) I Anillinus sp. “ Tennessee, Kings Saltpeter Cave ” J Anillinus virginiae (Virginia, Bath Co.) K Anillinus erwini (Virginia, Grayson Co.) L Serranillus sp. (North Carolina, Graham Co.) M Anillinus sp. “ South Carolina, Waldrop Stone ” (South Carolina, Pickens Co.) N Anillinus sp. “ Alabama, Horseshoe Cave sp. 1 ” O Anillinus loweae (North Carolina, Haywood Co.) P Anillinus sp. “ Kentucky, Hestand sp. 1 ” Q Anillinus indianae (Indiana, Lawrence Co.) R Anillinus cf. barberi (West Virginia, Greenbrier Co.) S Anillinus hirsutus (Alabama, Madison Co.). Scale bar: 1 mm." ;
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dc:description "Figure 10. SEM micrographs of male protarsi A Anillinus jancae, left protarsus, dorsal aspect B Anillinus mica, left protarsus, dorsal aspect C Anillinus montrex left protarsus, dorsal aspect D A. jancae, right protarsus, ventral aspect E Anillinus choestoea, right protarsus, ventral aspect F Anillinus dentatus, right protarsus, ventral aspect G Serranillus dunavani (Jeannel), right protarsus, ventral aspect H Anillinus murrayae Sokolov & Carlton, right protarsus, ventral aspect I A. montrex, right protarsus, ventral aspect. Scale bars: 0.1 mm." ;
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dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
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dc:description "Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." ;
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dc:description "Figure 34. Dorsal habitus of Anillinus species in the elongatus group and sinuaticollis group A Anillinus montrex B Anillinus arenicollis C Anillinus choestoea sp. nov. D Anillinus seneca sp. nov. E Anillinus mica sp. nov. F Anillinus micamicus sp. nov. Abdomens removed in B – F for DNA extraction. Scale bars: 1 mm." ;
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dc:description "Figure 10. SEM micrographs of male protarsi A Anillinus jancae, left protarsus, dorsal aspect B Anillinus mica, left protarsus, dorsal aspect C Anillinus montrex left protarsus, dorsal aspect D A. jancae, right protarsus, ventral aspect E Anillinus choestoea, right protarsus, ventral aspect F Anillinus dentatus, right protarsus, ventral aspect G Serranillus dunavani (Jeannel), right protarsus, ventral aspect H Anillinus murrayae Sokolov & Carlton, right protarsus, ventral aspect I A. montrex, right protarsus, ventral aspect. Scale bars: 0.1 mm." ;
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dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
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dc:description "Figure 23. Male genitalia of Anillinus dentatus (A – C) and Anillinus jancae (D – F). Median lobes in right lateral (C) and right dorsolateral (F) aspects. Left parameres (A, D) and right parameres (B, E) in left and right lateral aspects, respectively. Scale bars: 0.1 mm." ;
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dc:description "Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." ;
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dc:description "Figure 25. Anillini habitat in South Carolina A Waldrop Stone Falls, Pickens Co. (Anillinus mica, Anillinus micamicus, Anillinus murrayae, Serranillus dunavani) B Lower Whitewater River gorge, Oconee Co. (Anillinus cherokee, A. murrayae, Anillinus sp. “ South Carolina, Coon Branch ”, S. dunavani, Serranillus jeanneli, Serranillus sp. “ South Carolina, Coon Branch ” C Long Cane Creek, Abbeville Co. (Anillinus chandleri, Anillinus dentatus, Anillinus jancae, S. dunavani). The holotypes of A. mica and A. micamicus were collected under embedded rocks uphill of the wooden staircase in A. The holotype of A. jancae was collected in a pipe trap set on a hill just out of view in C." ;
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dc:description "Figure 43. Topology and distribution maps of “ quadrisetose clade ” species groups of Anillinus A collapsed maximum likelihood tree of “ quadrisetose clade ” Anillinus species, from 6 - gene concatenated core matrix, SBS values shown below nodes B distribution map of all known occurrences of Anillinus erwini and the valentinei group C distribution map of all known occurrences of the “ ESP + LSL ” clade and albrittonorum group D distribution map of all known occurrences of the “ ESP clade ” E distribution map of all known occurrences of the langdoni group and Anillinus sp. “ Tennessee, Kings Saltpeter Cave ” F distribution map of all known occurrences of the steevesi group G distribution map of all known occurrences of the loweae group. Data for distribution maps come from Harden (2024)." ;
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dc:creator "Harden, Curt W.; Caterino, Michael S." ;
dc:date "2024" ;
dc:title "Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)" ;
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dc:description "Figure 10. SEM micrographs of male protarsi A Anillinus jancae, left protarsus, dorsal aspect B Anillinus mica, left protarsus, dorsal aspect C Anillinus montrex left protarsus, dorsal aspect D A. jancae, right protarsus, ventral aspect E Anillinus choestoea, right protarsus, ventral aspect F Anillinus dentatus, right protarsus, ventral aspect G Serranillus dunavani (Jeannel), right protarsus, ventral aspect H Anillinus murrayae Sokolov & Carlton, right protarsus, ventral aspect I A. montrex, right protarsus, ventral aspect. Scale bars: 0.1 mm." ;
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dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
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dc:description "Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." ;
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dc:description "Figure 25. Anillini habitat in South Carolina A Waldrop Stone Falls, Pickens Co. (Anillinus mica, Anillinus micamicus, Anillinus murrayae, Serranillus dunavani) B Lower Whitewater River gorge, Oconee Co. (Anillinus cherokee, A. murrayae, Anillinus sp. “ South Carolina, Coon Branch ”, S. dunavani, Serranillus jeanneli, Serranillus sp. “ South Carolina, Coon Branch ” C Long Cane Creek, Abbeville Co. (Anillinus chandleri, Anillinus dentatus, Anillinus jancae, S. dunavani). The holotypes of A. mica and A. micamicus were collected under embedded rocks uphill of the wooden staircase in A. The holotype of A. jancae was collected in a pipe trap set on a hill just out of view in C." ;
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dc:description "Figure 30. Aedeagi of valentinei group Anillinus species A – C Anillinus castaneus sp. nov. D – F Anillinus cornelli G – I Anillinus simplex sp. nov. J – N Anillinus murrayae. Median lobes C, F, I in right dorsolateral aspect, median lobe K in right lateral aspect. Left A, D, G, J and right B, E, H, L – N parameres in left and right lateral aspects, respectively. Scale bar: 0.1 mm." ;
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dc:creator "Harden, Curt W.; Caterino, Michael S." ;
dc:date "2024" ;
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dwc:class "Insecta" ;
dwc:kingdom "Animalia" ;
dwc:order "Coleoptera" ;
dwc:phylum "Arthropoda" ;
dwc:rank "order" ;
trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Insecta> ;
a dwcFP:TaxonName .
<http://taxon-name.plazi.org/id/Animalia/Insecta>
dwc:class "Insecta" ;
dwc:kingdom "Animalia" ;
dwc:phylum "Arthropoda" ;
dwc:rank "class" ;
trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Arthropoda> ;
a dwcFP:TaxonName .
<http://taxon-name.plazi.org/id/Animalia/Arthropoda>
dwc:kingdom "Animalia" ;
dwc:phylum "Arthropoda" ;
dwc:rank "phylum" ;
trt:hasParentName <http://taxon-name.plazi.org/id/Animalia> ;
a dwcFP:TaxonName .
<http://taxon-name.plazi.org/id/Animalia>
dwc:kingdom "Animalia" ;
dwc:rank "kingdom" ;
a dwcFP:TaxonName .
<http://dx.doi.org/10.3897/zookeys.1209.125897.figure21>
dc:description "Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110360> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1209.125897.figure24>
dc:description "Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110363> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1209.125897.figure34>
dc:description "Figure 34. Dorsal habitus of Anillinus species in the elongatus group and sinuaticollis group A Anillinus montrex B Anillinus arenicollis C Anillinus choestoea sp. nov. D Anillinus seneca sp. nov. E Anillinus mica sp. nov. F Anillinus micamicus sp. nov. Abdomens removed in B – F for DNA extraction. Scale bars: 1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1110373> ;
a fabio:Figure .