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@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .
<http://treatment.plazi.org/id/2528091AFF88FFCF8BD3B81814840A33>
cito:cites <http://dx.doi.org/10.5281/zenodo.15214047>, <http://dx.doi.org/10.5281/zenodo.15214049>, <http://dx.doi.org/10.5281/zenodo.15214051>, <http://dx.doi.org/10.5281/zenodo.15214053>, <http://dx.doi.org/10.5281/zenodo.15214027>, <http://dx.doi.org/10.5281/zenodo.15214041>, <http://dx.doi.org/10.5281/zenodo.15214043>, <http://dx.doi.org/10.5281/zenodo.15214023> ;
dc:creator "Bohra, Sanath Chandra; Nguyen, Tan Van; Vogel, Gernot; Lalremsanga, Hmar Tlawmte; Biakzuala, Lal; Das, Madhurima; Warjri, Holiness; Thongni, Goldenstar; Poyarkov, Nikolay A.; Purkayastha, Jayaditya" ;
dc:title "Hebius gilhodesi Bohra, Nguyen, Vogel, Lalremsanga, Biakzuala, Das, Warjri, Thongni, Poyarkov & Purkayastha, 2025, comb. nov." ;
dwc:basisOfRecord <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF88FFCF8BD3B81814840A33.mc.1DE9B251FF8FFFF88BD3BFCB11480B24>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF88FFCF8BD3B81814840A33.mc.1DE9B251FF8FFFF88B83BC7F10EA0B19>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF88FFCF8BD3B81814840A33.mc.1DE9B251FF8FFFF88FD7BC7F15540B7D>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF88FFCF8BD3B81814840A33.mc.1DE9B251FF8FFFF88A7FBC5B16B50B7D>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF88FFCF8BD3B81814840A33.mc.1DE9B251FF8FFFF8899EBC5B10EE0B7D>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF88FFCF8BD3B81814840A33.mc.1DE9B251FF8FFFF88FD5BC5B157A0B51>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF88FFCF8BD3B81814840A33.mc.1DE9B251FF8FFFF88A61BC0716A70B51>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF88FFCF8BD3B81814840A33.mc.1DE9B251FF8FFFF8898CBC0710C60B51>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF88FFCF8BD3B81814840A33.mc.1DE9B251FF8FFFF88FEDBC0717F10BB5> ;
trt:citesTaxonName <http://taxon-name.plazi.org/id/Animalia/Natrix_gihodesi>, <http://taxon-name.plazi.org/id/Animalia/Natrix_khasiensis> ;
trt:definesTaxonConcept <http://taxon-concept.plazi.org/id/Animalia/Hebius_gilhodesi_(Bohra_1925> ;
trt:deprecates <http://taxon-concept.plazi.org/id/Animalia/Natrix_gilhodesi_Wall_1925> ;
trt:publishedIn <https://doi.org/10.11646/zootaxa.5604.4.3> ;
a trt:Treatment .
<https://doi.org/10.11646/zootaxa.5604.4.3>
bibo:endPage "504" ;
bibo:issue "4" ;
bibo:journal "Zootaxa" ;
bibo:pubDate "2025-03-17" ;
bibo:startPage "465" ;
bibo:volume "5604" ;
dc:creator "Bohra, Sanath Chandra; Nguyen, Tan Van; Vogel, Gernot; Lalremsanga, Hmar Tlawmte; Biakzuala, Lal; Das, Madhurima; Warjri, Holiness; Thongni, Goldenstar; Poyarkov, Nikolay A.; Purkayastha, Jayaditya" ;
dc:date "2025" ;
dc:title "Same but different: A systematic reassessment of the Hebius khasiensis Boulenger, 1890 (Reptilia: Squamata: Natricidae) species complex from the Indo-Burma biodiversity hotspot supports the revalidation of Natrix gilhodesi Wall, 1925 as a valid species" ;
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<http://taxon-concept.plazi.org/id/Animalia/Hebius_gilhodesi_(Bohra_1925>
dwc:authorityName "Bohra & Nguyen & Vogel & Lalremsanga & Biakzuala & Das & Warjri & Thongni & Poyarkov & Purkayastha" ;
dwc:authorityYear "2025" ;
dwc:baseAuthorityName "Bohra & Nguyen & Vogel & Lalremsanga & Biakzuala & Das & Warjri & Thongni & Poyarkov & Purkayastha" ;
dwc:baseAuthorityYear "1925" ;
dwc:class "Squamata" ;
dwc:family "Colubridae" ;
dwc:genus "Hebius" ;
dwc:kingdom "Animalia" ;
dwc:phylum "Chordata" ;
dwc:rank "species" ;
dwc:scientificNameAuthorship "(Bohra, Nguyen, Vogel, Lalremsanga, Biakzuala, Das, Warjri, Thongni, Poyarkov & Purkayastha, 1925)" ;
dwc:species "gilhodesi" ;
rdfs:seeAlso <https://www.catalogueoflife.org/data/taxon/8J9ML> ;
trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Hebius_gilhodesi> ;
a dwcFP:TaxonConcept .
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dwc:authorityName "Wall" ;
dwc:authorityYear "1925" ;
dwc:class "Squamata" ;
dwc:family "Colubridae" ;
dwc:genus "Natrix" ;
dwc:kingdom "Animalia" ;
dwc:phylum "Chordata" ;
dwc:rank "species" ;
dwc:scientificNameAuthorship "Wall, 1925" ;
dwc:species "gilhodesi" ;
trt:hasTaxonName <http://taxon-name.plazi.org/id/Animalia/Natrix_gilhodesi> ;
a dwcFP:TaxonConcept .
<http://taxon-name.plazi.org/id/Animalia/Hebius_gilhodesi>
dwc:class "Squamata" ;
dwc:family "Colubridae" ;
dwc:genus "Hebius" ;
dwc:kingdom "Animalia" ;
dwc:phylum "Chordata" ;
dwc:rank "species" ;
dwc:species "gilhodesi" ;
trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Hebius> ;
a dwcFP:TaxonName .
<http://taxon-name.plazi.org/id/Animalia/Hebius>
dwc:genus "Hebius" ;
dwc:kingdom "Animalia" ;
dwc:rank "genus" ;
trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Colubridae> ;
a dwcFP:TaxonName .
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dwc:family "Colubridae" ;
dwc:kingdom "Animalia" ;
dwc:rank "family" ;
trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Squamata> ;
a dwcFP:TaxonName .
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dwc:class "Squamata" ;
dwc:kingdom "Animalia" ;
dwc:rank "class" ;
trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Chordata> ;
a dwcFP:TaxonName .
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dwc:kingdom "Animalia" ;
dwc:phylum "Chordata" ;
dwc:rank "phylum" ;
trt:hasParentName <http://taxon-name.plazi.org/id/Animalia> ;
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<http://taxon-name.plazi.org/id/Animalia>
dwc:kingdom "Animalia" ;
dwc:rank "kingdom" ;
a dwcFP:TaxonName .
<http://taxon-name.plazi.org/id/Animalia/Natrix_gilhodesi>
dwc:class "Squamata" ;
dwc:family "Colubridae" ;
dwc:genus "Natrix" ;
dwc:kingdom "Animalia" ;
dwc:phylum "Chordata" ;
dwc:rank "species" ;
dwc:species "gilhodesi" ;
trt:hasParentName <http://taxon-name.plazi.org/id/Animalia/Natrix> ;
a dwcFP:TaxonName .
<http://taxon-name.plazi.org/id/Animalia/Natrix>
dwc:genus "Natrix" ;
dwc:kingdom "Animalia" ;
dwc:rank "genus" ;
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a dwcFP:TaxonName .
<http://dx.doi.org/10.5281/zenodo.15214047>
dc:description "FIGURE 10. Reproduced plates of sketches showing Hebius khasiensis (A and B– Sketch reproduced from Boulenger 1893b: Plate 13: Fig. 3) and Hebius gilhodesi comb. nov. (C and D– Sketch reproduced from original description i.e. Wall 1925a: Fig. 2)." ;
fabio:hasRepresentation <https://zenodo.org/record/15214047/files/figure.png> ;
a fabio:Figure .
<http://dx.doi.org/10.5281/zenodo.15214049>
dc:description "FIGURE 11. Holotype of Hebius gilhodesi comb. nov. (BMNH 1946.1.13.62, an adult male). A: dorsal view; B: ventral view; C: dorsal view of the head; D and E: lateral or side view of the head; F: ventral view of the head. Photographs by Gernot Vogel." ;
fabio:hasRepresentation <https://zenodo.org/record/15214049/files/figure.png> ;
a fabio:Figure .
<http://dx.doi.org/10.5281/zenodo.15214051>
dc:description "FIGURE 12. Dorsal view of paratypes of Hebius gilhodesi comb. nov. A: BMNH 1946.1.13.63 (adult male); B: BMNH 1925.4.2.10 (adult female); C: BMNH 1925.4.2.11 (adult male); D: BMNH 1925.4.2.12 (adult female); E: BMNH 1925.4.2.13 (adult male); F: BMNH 1925.4.2.14 (adult male); G: BMNH 1925.4.2.15 (adult female) and H: BMNH 1925.4.2.15A (adult female). Photographs by Gernot Vogel." ;
fabio:hasRepresentation <https://zenodo.org/record/15214051/files/figure.png> ;
a fabio:Figure .
<http://dx.doi.org/10.5281/zenodo.15214053>
dc:description "FIGURE 13. An uncollected adult specimen of Hebius gilhodesi comb. nov. from nearby Bhamo district, Kachin State, Myanmar. Note the presence of an indistinct, broken dorso-lateral streak on the nape. Photograph by K.C. Ouyang." ;
fabio:hasRepresentation <https://zenodo.org/record/15214053/files/figure.png> ;
a fabio:Figure .
<http://dx.doi.org/10.5281/zenodo.15214027>
dc:description "FIGURE 2. Maximum likelihood phylogeny showing phylogenetic relationships among members of the genus Hebius based on the Cytochrome b mitochondrial gene. Numbers at the nodes represent ultrafast bootstrap support. The species name is preceding by the NCBI accession number." ;
fabio:hasRepresentation <https://zenodo.org/record/15214027/files/figure.png> ;
a fabio:Figure .
<http://dx.doi.org/10.5281/zenodo.15214041>
dc:description "FIGURE 7. Comparison of the lateral regions of the head between Hebius khasiensis and Hebius gilhodesi comb. nov. A, B, C, D and E: Hebius gilhodesi comb. nov. while F, G, H, I and J: Hebius khasiensis. Note the strong differences between the two species in the dorso-lateral streak on the nape i.e. continuous in H. khasiensis versus broken into spots in H. gilhodesi comb. nov. Photographs by Gernot Vogel." ;
fabio:hasRepresentation <https://zenodo.org/record/15214041/files/figure.png> ;
a fabio:Figure .
<http://dx.doi.org/10.5281/zenodo.15214043>
dc:description "FIGURE 8. Comparisons of the dorsal scale rows on the anterior portion (OSR) of the body between Hebius khasiensis and Hebius gilhodesi comb. nov. A: Hebius khasiensis (very feebly keeled scales). B: Hebius gilhodesi comb. nov. (strongly keeled scales). Photographs by Gernot Vogel." ;
fabio:hasRepresentation <https://zenodo.org/record/15214043/files/figure.png> ;
a fabio:Figure .
<http://dx.doi.org/10.5281/zenodo.15214023>
dc:description "FIGURE 1. Map showing the known distribution of Hebius khasiensis (orange circles indicate distribution; orange square box indicates type locality) and Hebius gilhodesi comb. nov. (purple circles indicate distribution; purple square box indicates type locality); Inset: Map showing the location of northeast India and Myanmar (Gray box)." ;
fabio:hasRepresentation <https://zenodo.org/record/15214023/files/figure.png> ;
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dwc:catalogNumber "1946.1.13.62" ;
dwc:collectionCode "BMNH" ;
dwc:locality "Huton" ;
dwc:recordedBy "Reverend Father Gilhodes" ;
dwc:typeStatus "holotype" ;
dwc:verbatimElevation "1372" ;
dwc:verbatimLatitude "24.75859" ;
dwc:verbatimLongitude "97.41182" ;
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dwc:catalogNumber "1946.1.13.63" ;
dwc:collectionCode "BMNH" ;
dwc:locality "Huton" ;
dwc:recordedBy "Reverend Father Gilhodes" ;
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dwc:verbatimElevation "1372" ;
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dwc:verbatimLongitude "97.41182" ;
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dwc:catalogNumber "1925.4.2.10" ;
dwc:collectionCode "BMNH" ;
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dwc:catalogNumber "1925.4.2.11" ;
dwc:collectionCode "BMNH" ;
dwc:locality "Huton" ;
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dwc:catalogNumber "1925.4.2.12" ;
dwc:collectionCode "BMNH" ;
dwc:locality "Huton" ;
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dwc:typeStatus "paratype" ;
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dwc:catalogNumber "1925.4.2.13" ;
dwc:collectionCode "BMNH" ;
dwc:locality "Huton" ;
dwc:recordedBy "Reverend Father Gilhodes" ;
dwc:typeStatus "paratype" ;
dwc:verbatimElevation "1372" ;
dwc:verbatimLatitude "24.75859" ;
dwc:verbatimLongitude "97.41182" ;
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dwc:catalogNumber "1925.4.2.14" ;
dwc:collectionCode "BMNH" ;
dwc:locality "Huton" ;
dwc:recordedBy "Reverend Father Gilhodes" ;
dwc:typeStatus "paratype" ;
dwc:verbatimElevation "1372" ;
dwc:verbatimLatitude "24.75859" ;
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dwc:locality "Huton" ;
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dwc:locality "Huton" ;
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dwc:typeStatus "paratype" ;
dwc:verbatimElevation "1372" ;
dwc:verbatimLatitude "24.75859" ;
dwc:verbatimLongitude "97.41182" ;
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@prefix rdf: <http://www.w3.org/1999/02/22-rdf-syntax-ns#> .
@prefix rdfs: <http://www.w3.org/2000/01/rdf-schema#> .
@prefix bibo: <http://purl.org/ontology/bibo/> .
@prefix cito: <http://purl.org/spar/cito/> .
@prefix dc: <http://purl.org/dc/elements/1.1/> .
@prefix dwc: <http://rs.tdwg.org/dwc/terms/> .
@prefix dwcFP: <http://filteredpush.org/ontologies/oa/dwcFP#> .
@prefix fabio: <http://purl.org/spar/fabio/> .
@prefix trt: <http://plazi.org/vocab/treatment#> .
<http://treatment.plazi.org/id/2528091AFF92FFFF8BD3BBCD14840F06>
cito:cites <http://dx.doi.org/10.5281/zenodo.15214033>, <http://dx.doi.org/10.5281/zenodo.15214037>, <http://dx.doi.org/10.5281/zenodo.15214039>, <http://dx.doi.org/10.5281/zenodo.15214045>, <http://dx.doi.org/10.5281/zenodo.15214047>, <http://dx.doi.org/10.5281/zenodo.15214031>, <http://dx.doi.org/10.5281/zenodo.15214027>, <http://dx.doi.org/10.5281/zenodo.15214041>, <http://dx.doi.org/10.5281/zenodo.15214043>, <http://dx.doi.org/10.5281/zenodo.15214055>, <http://dx.doi.org/10.5281/zenodo.15214023> ;
dc:creator "Bohra, Sanath Chandra; Nguyen, Tan Van; Vogel, Gernot; Lalremsanga, Hmar Tlawmte; Biakzuala, Lal; Das, Madhurima; Warjri, Holiness; Thongni, Goldenstar; Poyarkov, Nikolay A.; Purkayastha, Jayaditya" ;
dc:title "Hebius khasiensis" ;
dwc:basisOfRecord <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF92FFFF8BD3BBCD14840F06.mc.1DE9B251FF91FFE68AD0BFC817E608C7>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF92FFFF8BD3BBCD14840F06.mc.1DE9B251FF91FFE688C3BFB510C508C7>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF92FFFF8BD3BBCD14840F06.mc.1DE9B251FF91FFE68FECBFB510780B3A>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF92FFFF8BD3BBCD14840F06.mc.1DE9B251FF91FFE68F6DBF9111390B1F>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF92FFFF8BD3BBCD14840F06.mc.1DE9B251FF91FFE68B83BC58102F0B57>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF92FFFF8BD3BBCD14840F06.mc.1DE9B251FF91FFE68B83BCE017440B8B>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF92FFFF8BD3BBCD14840F06.mc.1DE9B251FF91FFE68875BCE016770BEF>, <http://tb.plazi.org/GgServer/dwcaRecords/2528091AFF92FFFF8BD3BBCD14840F06.mc.1DE9B251FF91FFE68952BCCD15C30BC3> ;
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bibo:endPage "504" ;
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bibo:journal "Zootaxa" ;
bibo:pubDate "2025-03-17" ;
bibo:startPage "465" ;
bibo:volume "5604" ;
dc:creator "Bohra, Sanath Chandra; Nguyen, Tan Van; Vogel, Gernot; Lalremsanga, Hmar Tlawmte; Biakzuala, Lal; Das, Madhurima; Warjri, Holiness; Thongni, Goldenstar; Poyarkov, Nikolay A.; Purkayastha, Jayaditya" ;
dc:date "2025" ;
dc:title "Same but different: A systematic reassessment of the Hebius khasiensis Boulenger, 1890 (Reptilia: Squamata: Natricidae) species complex from the Indo-Burma biodiversity hotspot supports the revalidation of Natrix gilhodesi Wall, 1925 as a valid species" ;
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dc:description "FIGURE 4. Designated lectotype (an adult female) of Hebius khasiensis (BMNH 1946.1.12.80). A: dorsal view; B: ventral view; C and D: lateral or side view of the head; E: ventral view of the head. Photographs by Gernot Vogel." ;
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dc:description "FIGURE 5. Two out of the three paralectotypes of Hebius khasiensis. A (dorsal view) and B (ventral view): BMNH 1946.1.12.81 (adult male); C (dorsal view) and D (ventral view): BMNH 1946.1.12.82 (adult male). Photographs by Gernot Vogel." ;
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dc:description "FIGURE 6. One of the recently collected referred specimens of Hebius khasiensis (MZMU 3526, an adult female) from Mairang, Eastern West Khasi Hills, Meghalaya.A: dorsal view; B: ventral view; C: midbody showing strongly keeled scales on the dorsum and dark ventrolateral spots on the ventral edges; D: dorsal view of the head; E: lateral or side view of the head; F: ventral view of the head; Bottom: maxillary dentition (MZMU 3526). Photographs by Sanath Chandra Bohra." ;
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dc:description "FIGURE 9. Dorsal coloration of Hebius khasiensis in life A: MZMU 3526 (adult female from Mairang, Eastern West Khasi Hills, Meghalaya, India); B, C and D: uncollected adult specimens of Hebius khasiensis (unsexed individuals from Tamdil, Aizawl district, Mizoram, India). Photograph A was captured by Sanath Chandra Bohra. Photograph B was captured by Parag Shide. Photographs C and D were captured by Girish Choure." ;
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dc:description "FIGURE 10. Reproduced plates of sketches showing Hebius khasiensis (A and B– Sketch reproduced from Boulenger 1893b: Plate 13: Fig. 3) and Hebius gilhodesi comb. nov. (C and D– Sketch reproduced from original description i.e. Wall 1925a: Fig. 2)." ;
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dc:description "FIGURE 3. Scatterplot of the first two PCs in the principal component analysis (PCA) of the morphological data of Hebius khasiensis sensu stricto and Hebius gilhodesi comb. nov. superimposed by convex hull polygons. The morphological data used in the analysis are shown in Table 3." ;
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dc:description "FIGURE 2. Maximum likelihood phylogeny showing phylogenetic relationships among members of the genus Hebius based on the Cytochrome b mitochondrial gene. Numbers at the nodes represent ultrafast bootstrap support. The species name is preceding by the NCBI accession number." ;
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dc:description "FIGURE 7. Comparison of the lateral regions of the head between Hebius khasiensis and Hebius gilhodesi comb. nov. A, B, C, D and E: Hebius gilhodesi comb. nov. while F, G, H, I and J: Hebius khasiensis. Note the strong differences between the two species in the dorso-lateral streak on the nape i.e. continuous in H. khasiensis versus broken into spots in H. gilhodesi comb. nov. Photographs by Gernot Vogel." ;
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dc:description "FIGURE 8. Comparisons of the dorsal scale rows on the anterior portion (OSR) of the body between Hebius khasiensis and Hebius gilhodesi comb. nov. A: Hebius khasiensis (very feebly keeled scales). B: Hebius gilhodesi comb. nov. (strongly keeled scales). Photographs by Gernot Vogel." ;
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dc:description "FIGURE 14. An overview of the habitats where individuals of Hebius khasiensis were observed (Top photo: Sohra, East Khasi Hills, Meghalaya, India; Bottom photo: Mawkyrwat, South West Khasi Hills, Meghalaya, India). Photographs by Sanath Chandra Bohra." ;
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dc:description "FIGURE 1. Map showing the known distribution of Hebius khasiensis (orange circles indicate distribution; orange square box indicates type locality) and Hebius gilhodesi comb. nov. (purple circles indicate distribution; purple square box indicates type locality); Inset: Map showing the location of northeast India and Myanmar (Gray box)." ;
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