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dc:title "A review of the species groups of the Western Hemisphere Onthophagus Latreille (Coleoptera: Scarabaeidae: Scarabaeinae) using COI barcoding and gene trees" ;
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dc:description "FIG. 5. Onthophagus important characters marked with arrows (details discussed in the text). 1, O. atriglabrus; 2, O. orphnoides; 3, O. propraecellens; 4, O. genuinus; 5, O. andersoni; 6, O. crinitus; 7, O. gazellinus; 8, O. onthochromus; 9. O. landolti; 10, O. anthracinus; 11, O. championi; 12, O. cynomysi; 13, O. orpheus orpheus; 14, O. hoepfneri; 15, O. acuminatus; 16, O. incensus; 17, O. cyanellus; 18, O. sanpabloetlorum; 19, O. cartwrighti; 20, O. velutinus. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 8. Maximum-likelihood gene tree for the Western Hemisphere Onthophagus species. Bootstrap values were calculated using the rapid bootstrap algorithm of RaxML under the GTRCAT model with 10,000 replicates and run on the CIPRES portal. See Table 2 for an explanation of the specimen codes." ;
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dc:description "FIG. 5. Onthophagus important characters marked with arrows (details discussed in the text). 1, O. atriglabrus; 2, O. orphnoides; 3, O. propraecellens; 4, O. genuinus; 5, O. andersoni; 6, O. crinitus; 7, O. gazellinus; 8, O. onthochromus; 9. O. landolti; 10, O. anthracinus; 11, O. championi; 12, O. cynomysi; 13, O. orpheus orpheus; 14, O. hoepfneri; 15, O. acuminatus; 16, O. incensus; 17, O. cyanellus; 18, O. sanpabloetlorum; 19, O. cartwrighti; 20, O. velutinus. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 5. Onthophagus important characters marked with arrows (details discussed in the text). 1, O. atriglabrus; 2, O. orphnoides; 3, O. propraecellens; 4, O. genuinus; 5, O. andersoni; 6, O. crinitus; 7, O. gazellinus; 8, O. onthochromus; 9. O. landolti; 10, O. anthracinus; 11, O. championi; 12, O. cynomysi; 13, O. orpheus orpheus; 14, O. hoepfneri; 15, O. acuminatus; 16, O. incensus; 17, O. cyanellus; 18, O. sanpabloetlorum; 19, O. cartwrighti; 20, O. velutinus. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 5. Onthophagus important characters marked with arrows (details discussed in the text). 1, O. atriglabrus; 2, O. orphnoides; 3, O. propraecellens; 4, O. genuinus; 5, O. andersoni; 6, O. crinitus; 7, O. gazellinus; 8, O. onthochromus; 9. O. landolti; 10, O. anthracinus; 11, O. championi; 12, O. cynomysi; 13, O. orpheus orpheus; 14, O. hoepfneri; 15, O. acuminatus; 16, O. incensus; 17, O. cyanellus; 18, O. sanpabloetlorum; 19, O. cartwrighti; 20, O. velutinus. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 2. A summarized neighbor-joining tree for Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any group. The male metafemora of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 8. Maximum-likelihood gene tree for the Western Hemisphere Onthophagus species. Bootstrap values were calculated using the rapid bootstrap algorithm of RaxML under the GTRCAT model with 10,000 replicates and run on the CIPRES portal. See Table 2 for an explanation of the specimen codes." ;
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dc:description "FIG. 9. Maximum-likelihood gene tree for the Western Hemisphere Onthophagus species, partitioned by codon. An IQ-TREE analysis was performed using default parameters for the Invertebrate Mitochondrial Code. See Table 2 for an explanation of the specimen codes." ;
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dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 2. A summarized neighbor-joining tree for Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any group. The male metafemora of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 8. Maximum-likelihood gene tree for the Western Hemisphere Onthophagus species. Bootstrap values were calculated using the rapid bootstrap algorithm of RaxML under the GTRCAT model with 10,000 replicates and run on the CIPRES portal. See Table 2 for an explanation of the specimen codes." ;
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dc:description "FIG. 9. Maximum-likelihood gene tree for the Western Hemisphere Onthophagus species, partitioned by codon. An IQ-TREE analysis was performed using default parameters for the Invertebrate Mitochondrial Code. See Table 2 for an explanation of the specimen codes." ;
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View file

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dc:title "Onthophagus hecate" ;
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dc:title "A review of the species groups of the Western Hemisphere Onthophagus Latreille (Coleoptera: Scarabaeidae: Scarabaeinae) using COI barcoding and gene trees" ;
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dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 5. Onthophagus important characters marked with arrows (details discussed in the text). 1, O. atriglabrus; 2, O. orphnoides; 3, O. propraecellens; 4, O. genuinus; 5, O. andersoni; 6, O. crinitus; 7, O. gazellinus; 8, O. onthochromus; 9. O. landolti; 10, O. anthracinus; 11, O. championi; 12, O. cynomysi; 13, O. orpheus orpheus; 14, O. hoepfneri; 15, O. acuminatus; 16, O. incensus; 17, O. cyanellus; 18, O. sanpabloetlorum; 19, O. cartwrighti; 20, O. velutinus. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 6. Ratio of maximum width to length of metafemur, a larger number indicates a relatively wider femur. 1 O. curvicornis species group; 2 O. hircus species group; 3 O. nasutus species group; 4 O. mexicanus species group; 5 O. eulophus species group; 6 O. velutinus species group; 7 O. landolti species group; 8 O. hecate species group; 9 O. chevrolati species group; 10 O. crinitus species group; 11 O. gazellinus species group; 12 O. hoepfneri species group; 13 O. dicranius species group; O. clypeatus species group. Parts of the box plot: left end of the line, minimum value; right end of the line, maximum value; left side of the box, first quantile; right side of the box, fourth quantile; cross line in the box, median; X-shaped mark, average. See Table 2 for an explanation of the specimen codes." ;
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View file

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dc:description "FIG. 5. Onthophagus important characters marked with arrows (details discussed in the text). 1, O. atriglabrus; 2, O. orphnoides; 3, O. propraecellens; 4, O. genuinus; 5, O. andersoni; 6, O. crinitus; 7, O. gazellinus; 8, O. onthochromus; 9. O. landolti; 10, O. anthracinus; 11, O. championi; 12, O. cynomysi; 13, O. orpheus orpheus; 14, O. hoepfneri; 15, O. acuminatus; 16, O. incensus; 17, O. cyanellus; 18, O. sanpabloetlorum; 19, O. cartwrighti; 20, O. velutinus. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 2. A summarized neighbor-joining tree for Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any group. The male metafemora of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 8. Maximum-likelihood gene tree for the Western Hemisphere Onthophagus species. Bootstrap values were calculated using the rapid bootstrap algorithm of RaxML under the GTRCAT model with 10,000 replicates and run on the CIPRES portal. See Table 2 for an explanation of the specimen codes." ;
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dc:description "FIG. 9. Maximum-likelihood gene tree for the Western Hemisphere Onthophagus species, partitioned by codon. An IQ-TREE analysis was performed using default parameters for the Invertebrate Mitochondrial Code. See Table 2 for an explanation of the specimen codes." ;
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View file

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dc:title "Onthophagus gazellinus Bates 1887" ;
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dc:title "A review of the species groups of the Western Hemisphere Onthophagus Latreille (Coleoptera: Scarabaeidae: Scarabaeinae) using COI barcoding and gene trees" ;
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dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 2. A summarized neighbor-joining tree for Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any group. The male metafemora of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:title "Onthophagus curvicornis Latreille 1812" ;
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dc:title "A review of the species groups of the Western Hemisphere Onthophagus Latreille (Coleoptera: Scarabaeidae: Scarabaeinae) using COI barcoding and gene trees" ;
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dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 8. Maximum-likelihood gene tree for the Western Hemisphere Onthophagus species. Bootstrap values were calculated using the rapid bootstrap algorithm of RaxML under the GTRCAT model with 10,000 replicates and run on the CIPRES portal. See Table 2 for an explanation of the specimen codes." ;
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dc:description "FIG. 5. Onthophagus important characters marked with arrows (details discussed in the text). 1, O. atriglabrus; 2, O. orphnoides; 3, O. propraecellens; 4, O. genuinus; 5, O. andersoni; 6, O. crinitus; 7, O. gazellinus; 8, O. onthochromus; 9. O. landolti; 10, O. anthracinus; 11, O. championi; 12, O. cynomysi; 13, O. orpheus orpheus; 14, O. hoepfneri; 15, O. acuminatus; 16, O. incensus; 17, O. cyanellus; 18, O. sanpabloetlorum; 19, O. cartwrighti; 20, O. velutinus. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 2. A summarized neighbor-joining tree for Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any group. The male metafemora of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:creator "Kohlmann, Bert; Solís, Ángel" ;
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dc:description "FIG. 5. Onthophagus important characters marked with arrows (details discussed in the text). 1, O. atriglabrus; 2, O. orphnoides; 3, O. propraecellens; 4, O. genuinus; 5, O. andersoni; 6, O. crinitus; 7, O. gazellinus; 8, O. onthochromus; 9. O. landolti; 10, O. anthracinus; 11, O. championi; 12, O. cynomysi; 13, O. orpheus orpheus; 14, O. hoepfneri; 15, O. acuminatus; 16, O. incensus; 17, O. cyanellus; 18, O. sanpabloetlorum; 19, O. cartwrighti; 20, O. velutinus. See Table 7 for the author names of the species listed." ;
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<http://dx.doi.org/10.5281/zenodo.15213969>
dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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<http://dx.doi.org/10.5281/zenodo.15213984>
dc:description "FIG. 8. Maximum-likelihood gene tree for the Western Hemisphere Onthophagus species. Bootstrap values were calculated using the rapid bootstrap algorithm of RaxML under the GTRCAT model with 10,000 replicates and run on the CIPRES portal. See Table 2 for an explanation of the specimen codes." ;
fabio:hasRepresentation <https://zenodo.org/record/15213984/files/figure.png> ;
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<http://dx.doi.org/10.5281/zenodo.15213986>
dc:description "FIG. 9. Maximum-likelihood gene tree for the Western Hemisphere Onthophagus species, partitioned by codon. An IQ-TREE analysis was performed using default parameters for the Invertebrate Mitochondrial Code. See Table 2 for an explanation of the specimen codes." ;
fabio:hasRepresentation <https://zenodo.org/record/15213986/files/figure.png> ;
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View file

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dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 5. Onthophagus important characters marked with arrows (details discussed in the text). 1, O. atriglabrus; 2, O. orphnoides; 3, O. propraecellens; 4, O. genuinus; 5, O. andersoni; 6, O. crinitus; 7, O. gazellinus; 8, O. onthochromus; 9. O. landolti; 10, O. anthracinus; 11, O. championi; 12, O. cynomysi; 13, O. orpheus orpheus; 14, O. hoepfneri; 15, O. acuminatus; 16, O. incensus; 17, O. cyanellus; 18, O. sanpabloetlorum; 19, O. cartwrighti; 20, O. velutinus. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 2. A summarized neighbor-joining tree for Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any group. The male metafemora of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:title "Onthophagus nasutus Guerin-Meneville 1855" ;
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dc:title "A review of the species groups of the Western Hemisphere Onthophagus Latreille (Coleoptera: Scarabaeidae: Scarabaeinae) using COI barcoding and gene trees" ;
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dc:description "FIG. 5. Onthophagus important characters marked with arrows (details discussed in the text). 1, O. atriglabrus; 2, O. orphnoides; 3, O. propraecellens; 4, O. genuinus; 5, O. andersoni; 6, O. crinitus; 7, O. gazellinus; 8, O. onthochromus; 9. O. landolti; 10, O. anthracinus; 11, O. championi; 12, O. cynomysi; 13, O. orpheus orpheus; 14, O. hoepfneri; 15, O. acuminatus; 16, O. incensus; 17, O. cyanellus; 18, O. sanpabloetlorum; 19, O. cartwrighti; 20, O. velutinus. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 6. Ratio of maximum width to length of metafemur, a larger number indicates a relatively wider femur. 1 O. curvicornis species group; 2 O. hircus species group; 3 O. nasutus species group; 4 O. mexicanus species group; 5 O. eulophus species group; 6 O. velutinus species group; 7 O. landolti species group; 8 O. hecate species group; 9 O. chevrolati species group; 10 O. crinitus species group; 11 O. gazellinus species group; 12 O. hoepfneri species group; 13 O. dicranius species group; O. clypeatus species group. Parts of the box plot: left end of the line, minimum value; right end of the line, maximum value; left side of the box, first quantile; right side of the box, fourth quantile; cross line in the box, median; X-shaped mark, average. See Table 2 for an explanation of the specimen codes." ;
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dc:description "FIG. 2. A summarized neighbor-joining tree for Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any group. The male metafemora of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
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dc:description "FIG. 5. Onthophagus important characters marked with arrows (details discussed in the text). 1, O. atriglabrus; 2, O. orphnoides; 3, O. propraecellens; 4, O. genuinus; 5, O. andersoni; 6, O. crinitus; 7, O. gazellinus; 8, O. onthochromus; 9. O. landolti; 10, O. anthracinus; 11, O. championi; 12, O. cynomysi; 13, O. orpheus orpheus; 14, O. hoepfneri; 15, O. acuminatus; 16, O. incensus; 17, O. cyanellus; 18, O. sanpabloetlorum; 19, O. cartwrighti; 20, O. velutinus. See Table 7 for the author names of the species listed." ;
fabio:hasRepresentation <https://zenodo.org/record/15213978/files/figure.png> ;
a fabio:Figure .
<http://dx.doi.org/10.5281/zenodo.15213969>
dc:description "FIG. 1. Neighbor-joining tree for the Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any species group. The male protibiae of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
fabio:hasRepresentation <https://zenodo.org/record/15213969/files/figure.png> ;
a fabio:Figure .
<http://dx.doi.org/10.5281/zenodo.15213972>
dc:description "FIG. 2. A summarized neighbor-joining tree for Western Hemisphere Onthophagus species, based on K2P genetic distances comparisons between COI-5P mitochondrial barcoding sequences. Figs. 2325 belong to the O. eulophus species group. Figs. 44 and 63 are not assigned to any group. The male metafemora of the following species are depicted: 1, O. acuminatus; 2, O. batesi; 3, O. incensus; 4, O. nitidor; 5, O. stockwelli; 6, O. striatulus striatulus; 7, O. tristis; 8, O. basicarinatus; 9, O. hircus; 10, O. marginicollis; 11, O. nyctopus; 12, O. onorei; 13, O. osculati; 14, O. ptox; 15, O. ranunculus; 16, O. atriglabrus; 17, O. genuinus; 18, O. rostratus; 19, O. sharpi; 20, O. tapirus; 21, O. villanuevai; 22, O. violetae; 23, O. anewtoni; 24, O. arnetti; 25, O. browni; 26, O. championi; 27, O. guatemalensis; 28, O. mexicanus; 29, O. cartwrighti; 30, O. velutinus; 31, O. alluvius; 32, O. altivagans; 33, O. anthracinus; 34, O. brachypterus; 35, O. chryses; 36, O. landolti; 37, O. lecontei; 38, O. schaefferi; 39, O. concinnus; 40, O. cynomysi; 41, O. hecate hecate; 42, O. medorensis; 43, O. orpheus orpheus; 44, O. polyphemi sparsisetosus; 45, O. chevrolati; 46, O. cyanellus, 47, O. hippopotamus; 48, O. oaxacanus; 49, O. reyesi; 50, O. salvadorensis; 51, O. sanpabloetlorum; 52, O. skelleyi; 53, O. subtropicus; 54, O. totonicapanus; 55, O. vespertilio; 56, O. crinitus; 57, O. gazellinus; 58, O. onthochromus; 59, O. aciculatulus; 60, O. coscineus; 61, O. hoepfneri; 62, O. igualensis; 63, O. lebasi; 64, O. pennsylvanicus; 65, O. subanaeus; 66, tuberculifrons; 67, O. barreti; 68, O.breviconus; 69, O. contrapositus; 70, O. dicranius; 71, O. dorsipilulus; 72, O. humboldti; 73, O. kohlmanni; 74, O. micropterus; 75, O. neomirabilis; 76, O. orphnoides; 77, O. petenensis; 78, O. subcancer; 79, O. andersoni; 80, O. belorhinus; 81, O. grataehelenae; 82, O. limonensis; 83, O. maya; 84, O. praecellens; 85, O. propraecellens; 86, O. rhinolophus; 87, O. rhinophyllus; 88, undescribed species of Onthophagus; 89, O. viridivinosus; 90, O. xanthomerus. See Table 2 for an explanation of the specimen codes. See Table 7 for the author names of the species listed." ;
fabio:hasRepresentation <https://zenodo.org/record/15213972/files/figure.png> ;
a fabio:Figure .