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cito:cites <http://dx.doi.org/10.3897/zookeys.1207.117225.figure4>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure6>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure8>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure16>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure17>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure18>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure23>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure26>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure33>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure34>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure29>, <http://dx.doi.org/10.3897/zookeys.1207.117225.figure27> ;
dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:title "Aphodius (Liothorax) krelli Angus, Maté, Angus & Král, 2024, sp. nov." ;
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bibo:endPage "299" ;
bibo:journal "ZooKeys" ;
bibo:pubDate "2024-07-22" ;
bibo:startPage "205" ;
bibo:volume "1207" ;
dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:date "2024" ;
dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
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dwc:authorityName "Angus & Maté & Angus & Král" ;
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dwc:class "Insecta" ;
dwc:family "Scarabaeidae" ;
dwc:genus "Aphodius" ;
dwc:kingdom "Animalia" ;
dwc:order "Coleoptera" ;
dwc:phylum "Arthropoda" ;
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dwc:class "Insecta" ;
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dwc:genus "Aphodius" ;
dwc:kingdom "Animalia" ;
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dc:description "Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." ;
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dc:description "Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097465> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure8>
dc:description "Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097467> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure16>
dc:description "Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097478> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure17>
dc:description "Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097480> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure18>
dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097482> ;
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<http://dx.doi.org/10.3897/zookeys.1207.117225.figure23>
dc:description "Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097492> ;
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dc:description "Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097499> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure33>
dc:description "Figure 33. Mitotic chromosomes arranged as karyotypes a – d A. bameuli sp. nov., ♂ paratypes, FR, Corsica, by the Lac de Melo. Autosome 9 polymorphic in a, b homozygous submetacentric in c, d. a, c plain b and d the same nuclei C-banded e – l A. krelli sp. nov., types, IT, Sardinia e, f paratype ♂ with the heterochromatic long arm of autosome 1 weakly developed e plain f the same nucleus C-banded g, h holotype with the heterochromatic long arm of autosome 1 fully developed g plain h the same nucleus C-banded i, j paratype ♂ with both replicates of autosome 7 lacking the heterochromatic long arm i plain j the same nucleus C-banded k, l paratype ♀ with autosome 9 heterozygous for an inversion polymorphism k plain l the same nucleus C-banded m – p A. wilsonae SP m, n holotype ♂ m plain n the same nucleus C-banded o, p paratype ♀ from Balneario de Corconte o plain p the same nucleus C-banded. Scale bar: 5 μm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097522> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure34>
dc:description "Figure 34. Meiotic chromosomes, metaphase I from testis a – d A. niger, GB, Hampshire, New Forest, White Moor a, c plain b, d the same nuclei C-banded e – g A. muscorum, HU, Hortobágyi e plain f, g the same nucleus f plain g C-banded h A. bameuli paratype, FR, Corsica, by Lac de Melo, plain i, j A. krelli IT, Sardinia, paratype i plain j the same nucleus C-banded k, l A. wilsonae, SP, Areños k plain l the same nucleus C-banded. The sex bivalent is indicated by an arrow. Scale bar: 5 μm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097526> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure29>
dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097509> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure27>
dc:description "Figure 27. Habitat pictures a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. niger, SW, Tullgarn d A. krelli sp. nov., IT, Sardinia, Badde Salighes e A. bameuli sp. nov., FR, Corsica, Haute-Corse, Lac de Melo f A. wilsonae, SP, Cantabria, Corconte g A. bellumgerens sp. nov., IT, Sicily, Nebrodi." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097502> ;
a fabio:Figure .
<http://tb.plazi.org/GgServer/dwcaRecords/121D8F6875C25860934C05AC5878D13E.mc.D51054BB0BB6093B82C6884CA8F1397D>
dwc:collectionCode "NHMUK" ;
dwc:eventDate "2012-04-03" ;
dwc:locality "Badde Salighes" ;
dwc:recordedBy "R. B. & E. M. Angus." ;
dwc:typeStatus "holotype" ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:title "Aphodius (Liothorax) rusakovi Gusakov 2004" ;
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bibo:endPage "299" ;
bibo:journal "ZooKeys" ;
bibo:pubDate "2024-07-22" ;
bibo:startPage "205" ;
bibo:volume "1207" ;
dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:date "2024" ;
dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
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dwc:authority "Gusakov, 2004" ;
dwc:authorityName "Gusakov" ;
dwc:authorityYear "2004" ;
dwc:class "Insecta" ;
dwc:family "Scarabaeidae" ;
dwc:genus "Aphodius" ;
dwc:kingdom "Animalia" ;
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dc:description "Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." ;
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dc:description "Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097463> ;
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dc:description "Figure 14. Aphodius plagiatus group, pronotal bases 2 a A. discoides, TR b A. kraatzi, SK c L. rusakovi, paratype, RU, Volgograd oblast’." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097474> ;
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dc:description "Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." ;
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dc:description "Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097486> ;
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dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
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dc:description "Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." ;
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dc:description "Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097499> ;
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dc:description "Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097466> ;
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dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097509> ;
a fabio:Figure .

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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:title "Aphodius (Liothorax) alberti Angus, Maté, Angus & Král, 2024, sp. nov." ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:date "2024" ;
dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
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dc:description "Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." ;
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dc:description "Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." ;
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dc:description "Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097465> ;
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dc:description "Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097467> ;
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dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097482> ;
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dc:description "Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097486> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure21>
dc:description "Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097488> ;
a fabio:Figure .
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dc:description "Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097494> ;
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dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097509> ;
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dc:description "Figure 28. Habitat pictures a A. plagiatus, RU, Pavlosk Park b A. rodrigoi sp. nov., SP, Toledo, Quero c A. p. sinoplagiatus, CN, Tibet, Ganca d A. kraatzi, RU, Astrakhan oblast’ e A. alberti sp. nov., TU, Ovit Dağı (photograph Copyright Yunus Yasuv) f A. felix sp. nov., IT, Campo Felice." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097505> ;
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dwc:collectionCode "NHMUK" ;
dwc:eventDate "1992-06-18" ;
dwc:locality "Ovitdagi" ;
dwc:recordedBy "A. Ballerio." ;
dwc:typeStatus "holotype" ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:title "Aphodius (Liothorax) chellala Angus, Maté, Angus & Král, 2024, sp. nov." ;
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bibo:volume "1207" ;
dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:date "2024" ;
dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
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dc:description "Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." ;
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dc:description "Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." ;
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dc:description "Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097466> ;
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dc:description "Figure 13. Aphodius plagiatus group, pronotal bases 1 a, b A. plagiatus a GB, Norfolk b MG c A. rutilipennis, CY d A. cf. rutilipennis, GR, Thessaloniki e A. chellala sp. nov., paratype, AG." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097473> ;
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dc:description "Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." ;
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dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
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dc:description "Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." ;
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dc:description "Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." ;
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dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
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dc:creator "Aung, Sein Lai Lai; Liu, Feng-Yin; Gou, Ya-Nan; Nwe, Zin Mar; Yu, Zhi-He; Deng, Jian-Xin" ;
dc:title "Alternaria momordicae S. L. L. Aung & J. X. Deng 2024, sp. nov." ;
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dc:date "2024" ;
dc:title "Morphological and phylogenetic analyses reveal two new Alternaria species (Pleosporales, Pleosporaceae) in Alternaria section from Cucurbitaceae plants in China" ;
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dc:description "Figure 3. Alternaria momordicae sp. nov. (ex-type YZU 161378) A, B seven-day-old culture on PDA C, D conidiophores and conidia on PCA and V 8 A, respectively E, F conidia on PCA and V 8 A, respectively. Scale bars: 25 μm (E, F); 50 μm (C, D)." ;
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dc:description "Figure 1. Phylogenetic tree of the Alternaria species most related to the new taxa based on maximum likelihood analysis using the combined gene sequences of ITS, GAPDH, TEF 1, RPB 2, Alt a 1, EndoPG and OPA 10-2 which rooted with Alternaria alternantherae (CBS 124392) from sect. Alternantherae. The Bayesian posterior probabilities > 0.60 (PP) and bootstrap support values > 60 (BS) are given at the nodes (PP / BS). The novel species are highlighted in bold. Ex-type isolates are marked with a superscript T and Representative isolates are marked with a superscript R." ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:title "Aphodius (Liothorax) wilsonae Mate" ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:date "2024" ;
dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
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dc:description "Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." ;
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dc:description "Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." ;
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dc:description "Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." ;
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dc:description "Figure 9. a – i epipharynxes a – c A. niger a SV, Tullgarn b, c GB, Hampshire, New Forest d A. niger?, CZ, Hradec Králové e A. muscorum, HU, Hortobagyi f, g A. bellumgerens sp. nov., IT Piano Battaglia, paratypes h, i A. wilsonae paratypes, SP h El Vellón i Manzanares el Real. Scale bar: 0.5 mm." ;
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dc:description "Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." ;
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dc:description "Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." ;
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dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
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dc:description "Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." ;
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dc:description "Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." ;
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dc:description "Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." ;
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dc:description "Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." ;
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dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
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dc:description "Figure 27. Habitat pictures a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. niger, SW, Tullgarn d A. krelli sp. nov., IT, Sardinia, Badde Salighes e A. bameuli sp. nov., FR, Corsica, Haute-Corse, Lac de Melo f A. wilsonae, SP, Cantabria, Corconte g A. bellumgerens sp. nov., IT, Sicily, Nebrodi." ;
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dwc:locality "Balneario de Corconte" ;
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dwc:verbatimLatitude "40.031" ;
dwc:verbatimLongitude "-3.884" ;
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dc:creator "Valentich-Scott, Paul; Griffiths, Charles; Landschoff, Jannes; Li, Ruiqi; Li, Jingchun" ;
dc:title "Brachiomya ducentiunus Valentich-Scott, Griffiths, Landschoff, Li & Li, 2024, sp. nov." ;
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dc:title "Bivalves of superfamily Galeommatoidea (Mollusca, Bivalvia) from western South Africa, with observations on commensal relationships and habitats" ;
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dc:description "Figure 1. Brachiomya ducentiunus sp. nov. A, B holotype, SBMNH 713162, length 2.50 mm, height 1.75 mm A exterior of right valve B exterior of left valve C – H paratypes, SBMNH 666970, crystallization on exterior and interior of shell is from dried bleach, not a sculpture element C exterior of right valve, length 2.17 mm, height 1.38 mm D exterior of left valve, length 2.40 mm, height 1.56 mm E interior of left valve, length 1.87 mm, height 1.19 mm F interior of right valve, length 2.42 mm, height 1.67 mm G hinge of right valve H hinge of left valve." ;
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dc:description "Figure 2. Brachiomya ducentiunus new species, living animals A, B crawling on hard substrate C overview of specimens crawling on the aboral surface of urchin Spatagobrissus mirabilis D crawling on urchin spine with mantle and foot extended E, F extended mantle between urchin spines." ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:title "Aphodius (Liothorax) bameuli Angus, Maté, Angus & Král, 2024, sp. nov." ;
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dc:date "2024" ;
dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
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dc:description "Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." ;
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dc:description "Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097465> ;
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dc:description "Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097467> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure16>
dc:description "Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." ;
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dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
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dc:description "Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097488> ;
a fabio:Figure .
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dc:description "Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." ;
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dc:description "Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097494> ;
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dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097509> ;
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dc:description "Figure 27. Habitat pictures a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. niger, SW, Tullgarn d A. krelli sp. nov., IT, Sardinia, Badde Salighes e A. bameuli sp. nov., FR, Corsica, Haute-Corse, Lac de Melo f A. wilsonae, SP, Cantabria, Corconte g A. bellumgerens sp. nov., IT, Sicily, Nebrodi." ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:title "Aphodius (Liothorax) isikdagensis" ;
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bibo:volume "1207" ;
dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:date "2024" ;
dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
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dc:description "Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." ;
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dc:description "Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." ;
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dc:description "Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." ;
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dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
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dc:description "Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." ;
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dc:description "Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." ;
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dc:description "Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" ;
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dc:description "Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." ;
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dc:description "Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." ;
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dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:title "Aphodius (Liothorax) plagiatus subsp. plagiatus" ;
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bibo:startPage "205" ;
bibo:volume "1207" ;
dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:date "2024" ;
dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
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dc:description "Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." ;
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dc:description "Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097463> ;
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dc:description "Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." ;
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dc:description "Figure 13. Aphodius plagiatus group, pronotal bases 1 a, b A. plagiatus a GB, Norfolk b MG c A. rutilipennis, CY d A. cf. rutilipennis, GR, Thessaloniki e A. chellala sp. nov., paratype, AG." ;
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dc:description "Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." ;
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dc:description "Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." ;
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dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
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dc:description "Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." ;
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dc:description "Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." ;
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dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
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dc:title "Aphodius (Liothorax) plagiatus subsp. sinoplagiatus Angus, Maté, Angus & Král, 2024, subsp. nov." ;
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dc:description "Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." ;
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dc:description "Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." ;
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dc:description "Figure 30. Mitotic chromosomes, arranged as karyotypes a, b A. plagiatus plagiatus, ♂, GB, Dorset, Studland a plain (Giemsa stained) b the same nucleus C-banded c A. plagiatus sinoplagiatus, ♂, CH, Qinghai, Gangca, plain d, e A. rutilipennis, ♂, CY, Zakaki Marshes d plain e C-banded f – h A. kraatzi ♀, SK f, g nucleus without B-chromosomes f plain g C-banded h, i nucleus with 1 B-chromosome h plain i C-banded. Scale bar: 5 μm." ;
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dc:description "Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." ;
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dc:description "Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." ;
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dc:description "Figure 15. Aphodius niger group, pronotal bases 1 a A. niger, Sweden, Södermanland, Tullgarn b A. cf. niger, CZ, České Budějovice c A. muscorum, HU, Hortobágyi d A. felix sp. nov., paratype." ;
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dc:description "Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." ;
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dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
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dc:description "Figure 19. Elytral apices a A. (L.) niger, SV, Södermanland, Tullgarn b A. (Nialus) varians, FR, left elytron c A. (Calamosternus) granarius, CY, right elytron d A. (C.) hyxos, CY, left elytron e A. (Agoliinus) lapponum, GB, Cumbria, both elytra f A. (Agrilinus) constans, IT, Sardinia, left elytron. Scale bar: 1 mm." ;
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dc:description "Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." ;
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dc:description "Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." ;
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dc:description "Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." ;
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dc:description "Figure 25. Aedeagal encasements (9 th abdominal sternite) a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. felix sp. nov., IT, Campo Felice d A. alberti sp. nov., TU, Ovit Dağı e A. bellumgerens sp. nov., IT, Sicily, Piano Battaglia f A. bameuli sp. nov., FR, Corsica, Haute-Corse g A. wilsonae, SP, Cantabria, Corconte h A. kraatzi, RU, Chita Reg. i A. plagiatus, UK, Norfolk, Hunstanton j A. plagiatus, CN, Beijing k A. rodrigoi sp. nov., SP, Madrid, Aranjuez l A. chellala sp. nov., AG, Chellala, paratype m A. rutilipennis sp. nov., CY, Konakli, paratype n A. discoides sp. nov., paratype. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097496> ;
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dc:description "Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." ;
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dc:description "Figure 31. Mitotic chromosomes, arranged as karyotypes a, b A. niger, ♂, SV, Tullgarn a plain b the same nucleus C-banded (1 replicate of autosome 5 missing) c A. niger ♀, SV, Tullgarn, C-banded d, e A. niger, ♂♂, GB, Hampshire, New Forest, C-banded e with 3 B-chromosomes f, g A. muscorum, ♂, HU, Hortobágyi f plain g the same nucleus C-banded h A. muscorum, ♂, HU, Hortobágyi, C-banded i, j A. muscorum, ♀, HU, Kisújszállás i plain j C-banded k, l A. muscorum, ♀, HU, Kisújszállás k plain l the same nucleus C-banded. Scale bar: 5 μm" ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097515> ;
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dc:description "Figure 34. Meiotic chromosomes, metaphase I from testis a – d A. niger, GB, Hampshire, New Forest, White Moor a, c plain b, d the same nuclei C-banded e – g A. muscorum, HU, Hortobágyi e plain f, g the same nucleus f plain g C-banded h A. bameuli paratype, FR, Corsica, by Lac de Melo, plain i, j A. krelli IT, Sardinia, paratype i plain j the same nucleus C-banded k, l A. wilsonae, SP, Areños k plain l the same nucleus C-banded. The sex bivalent is indicated by an arrow. Scale bar: 5 μm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097526> ;
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dc:description "Figure 27. Habitat pictures a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. niger, SW, Tullgarn d A. krelli sp. nov., IT, Sardinia, Badde Salighes e A. bameuli sp. nov., FR, Corsica, Haute-Corse, Lac de Melo f A. wilsonae, SP, Cantabria, Corconte g A. bellumgerens sp. nov., IT, Sicily, Nebrodi." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097502> ;
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dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:title "Aphodius (Liothorax) rodrigoi Angus, Maté, Angus & Král, 2024, sp. nov." ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:date "2024" ;
dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
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dc:description "Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." ;
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dc:description "Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." ;
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dc:description "Figure 11. Aphodius plagiatus group, maxillae a A. rusakovi, KZ, Syr-Darja b, c A. kraatzi b CZ c RU d A. plagiatus plagiatus, GB, Norfolk e A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca f A. rodrigoi sp. nov. paratype SP Madrid, Aranjuez g A chellala, paratype, AG, Chellala h A. rutilipennis CY i A. discoides, TR, Karacadag. Scale bar: 0.5 mm." ;
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dc:description "Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." ;
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dc:description "Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." ;
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dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
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dc:description "Figure 28. Habitat pictures a A. plagiatus, RU, Pavlosk Park b A. rodrigoi sp. nov., SP, Toledo, Quero c A. p. sinoplagiatus, CN, Tibet, Ganca d A. kraatzi, RU, Astrakhan oblast’ e A. alberti sp. nov., TU, Ovit Dağı (photograph Copyright Yunus Yasuv) f A. felix sp. nov., IT, Campo Felice." ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:title "Aphodius (Liothorax) bellumgerens Angus, Maté, Angus & Král, 2024, sp. nov." ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:date "2024" ;
dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
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dc:description "Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." ;
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dc:description "Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097465> ;
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dc:description "Figure 9. a – i epipharynxes a – c A. niger a SV, Tullgarn b, c GB, Hampshire, New Forest d A. niger?, CZ, Hradec Králové e A. muscorum, HU, Hortobagyi f, g A. bellumgerens sp. nov., IT Piano Battaglia, paratypes h, i A. wilsonae paratypes, SP h El Vellón i Manzanares el Real. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097468> ;
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dc:description "Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097478> ;
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dc:description "Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." ;
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dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
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dc:description "Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." ;
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dc:description "Figure 24. A. niger group aedeagi, cleared preparations mounted in DMHF on slides a – g, m – o dorsoventral view h – l, p right lateral view a A. niger, SV, Södermanland, Tullgarn b A. bameuli sp. nov., paratype c A. isikdagensis, paratype, TR, Ḉamildere d A. krelli sp. nov., paratype, IT, Sardinia e – f A. alberti sp. nov., TR, Rize, Ovitdag e paratype, after squashing f the same specimen, before squashing g, h, j A. alberti sp. nov., different paratypes i A. alberti sp. nov., paratype, paramere k A. isikdagensis?, TR, Artvin l A. alberti?, Armenia m A. bellumgerens sp. nov., paratype, IT, Sicily, Piano Battaglia n A. bellumgerens, paratype IT, Sicily, Nebrodi o A. wilsonae, paratype, SP, Manzanares el Real p A. muscorum, HU, Hortobági. Scale bar: 1 mm." ;
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dc:description "Figure 25. Aedeagal encasements (9 th abdominal sternite) a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. felix sp. nov., IT, Campo Felice d A. alberti sp. nov., TU, Ovit Dağı e A. bellumgerens sp. nov., IT, Sicily, Piano Battaglia f A. bameuli sp. nov., FR, Corsica, Haute-Corse g A. wilsonae, SP, Cantabria, Corconte h A. kraatzi, RU, Chita Reg. i A. plagiatus, UK, Norfolk, Hunstanton j A. plagiatus, CN, Beijing k A. rodrigoi sp. nov., SP, Madrid, Aranjuez l A. chellala sp. nov., AG, Chellala, paratype m A. rutilipennis sp. nov., CY, Konakli, paratype n A. discoides sp. nov., paratype. Scale bar: 0.5 mm." ;
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dc:description "Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." ;
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dc:description "Figure 32. Mitotic chromosomes arranged as karyotypes a – e A. felix sp. nov. IT, Abruzzo. Campo Felice a, b holotype ♂ a plain b the same nucleus C-banded c holotype ♂ C-banded, autosome 8 polymorphic, 9 subacrocentric d paratype ♂, C-banded, autosomes 8 and 9 metacentric e paratype ♀ f – i A. bellumgerens sp. nov., paratypes, IT, Sicily f, g A. bellumgerens sp. nov., ♂, paratype f plain g C-banded h, i A. bellumgerens sp. nov., ♀, paratype h plain i C-banded. Scale bar: 5 μm" ;
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dc:description "Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." ;
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dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
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dc:description "Figure 27. Habitat pictures a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. niger, SW, Tullgarn d A. krelli sp. nov., IT, Sardinia, Badde Salighes e A. bameuli sp. nov., FR, Corsica, Haute-Corse, Lac de Melo f A. wilsonae, SP, Cantabria, Corconte g A. bellumgerens sp. nov., IT, Sicily, Nebrodi." ;
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dc:creator "Aung, Sein Lai Lai; Liu, Feng-Yin; Gou, Ya-Nan; Nwe, Zin Mar; Yu, Zhi-He; Deng, Jian-Xin" ;
dc:title "Alternaria jingzhouensis S. L. L. Aung & J. X. Deng 2024, sp. nov." ;
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bibo:startPage "125" ;
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dc:date "2024" ;
dc:title "Morphological and phylogenetic analyses reveal two new Alternaria species (Pleosporales, Pleosporaceae) in Alternaria section from Cucurbitaceae plants in China" ;
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dc:description "Figure 2. Alternaria jingzhouensis sp. nov. (ex-type YZU 221144) A, B seven-day-old culture on PDA C, D conidiophores and conidia on PCA and V 8 A, respectively E, F conidia on PCA and V 8 A, respectively. Scale bars: 25 μm (E, F); 50 μm (C, D)." ;
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dc:description "Figure 1. Phylogenetic tree of the Alternaria species most related to the new taxa based on maximum likelihood analysis using the combined gene sequences of ITS, GAPDH, TEF 1, RPB 2, Alt a 1, EndoPG and OPA 10-2 which rooted with Alternaria alternantherae (CBS 124392) from sect. Alternantherae. The Bayesian posterior probabilities > 0.60 (PP) and bootstrap support values > 60 (BS) are given at the nodes (PP / BS). The novel species are highlighted in bold. Ex-type isolates are marked with a superscript T and Representative isolates are marked with a superscript R." ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:title "Aphodius (Liothorax) felix Angus, Maté, Angus & Král, 2024, sp. nov." ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:date "2024" ;
dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
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dc:description "Figure 4. a – g whole beetles a A. niger, SV, Tullgarn b A. muscorum, holotype HU, Vörös tó c A. felix sp. nov., holotype, IT, Campo Felice d A. bameuli sp. nov., holotype, Corsica, by Lac de Melo e A. krelli sp. nov., holotype, IT, Sardinia f A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia g A. wilsonae, holotype, SP, Provincia de Burgos, Balneario de Corconte. Scale bar: 1 mm." ;
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dc:description "Figure 6. a – l A. niger group, heads and pronota a, b A. isikdagensis, paratypes, TR Ḉamildere a ♂ b ♀ b ’ detail of right lateral part of frons and gena, different orientation b ” detail of right posterior of pronotum, different orientation c, d A. alberti sp. nov., TR, Rize, Ovitdagi c holotype d ♀ paratype e A. niger, GB, Hampshire, New Forest f A. muscorum, HU, Hortobágyi g A. bameuli sp. nov., holotype, FR, Corsica, by Lac de Melo h A. krelli sp. nov., holotype, IT, Sardinia i A. felix sp. nov., holotype, IT, Campo Felice j A. bellumgerens sp. nov., holotype, IT, Sicily, Piano Battaglia k, l A. wilsonae, SP k holotype, Provincia de Burgos, Balneario de Corconte l paratype, Provincia de Madrid, Manzanares el Real. Scale bar: 1 mm." ;
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dc:description "Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" ;
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dc:description "Figure 15. Aphodius niger group, pronotal bases 1 a A. niger, Sweden, Södermanland, Tullgarn b A. cf. niger, CZ, České Budějovice c A. muscorum, HU, Hortobágyi d A. felix sp. nov., paratype." ;
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dc:description "Figure 16. Aphodius niger group, pronotal bases 2 a A. bameuli sp. nov., paratype, FR, Corsica b A. krelli sp. nov., paratype, IT, Sardinia c, d A. wilsonae, paratypes, SP, Manzanares el Real e A. isikdagensis, paratype, TR, Ḉamildere." ;
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dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
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dc:description "Figure 21. Metaventrites, A. niger group a – d A. niger a, c, d SV, Södermanland, Tullgarn b GB, Hampshire, New Forest a, b ♂ c, d ♀ e, g – j A. muscorum e holotype ♀, HU, Vörös-tó f A. felix sp. nov., paratype ♂, IT, Campo Felice g ♂, HU, Hortobágyi h, i ♀♀, HU, Kisujszallas j ♀, HU, Josvafo k A. felix sp. nov. paratype ♀, IT, Campo Felice l, m A. bellumgerens sp. nov., paratypes, IT, Sicily, Piano Battaglia l ♂ m ♀ n, o A. alberti sp. nov., paratypes, TR, Rize, Ovitdag n ♂, o ♀ p – r A. bameuli sp. nov., paratypes, FR, Corsica p ♂ q, r ♀ s, t A. krelli sp. nov., paratypes, IT, Sardinia s ♂ t ♀ u – w A. wilsonae, paratypes u, v SP, Manzanares el Real u ♂ v ♀ w ♀, SP, El Vellon x, y A. isikdagensis, paratypes, TR, Ḉamildere x ♂ y ♀. Scale bar: 0.5 mm." ;
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dc:description "Figure 23. A. niger group, aedeagi with inflated endophalli, stacked photographs and SEM images a – e, g – w right lateral view f viewed from above a – c A. niger, SV, Södermanland, Tullgarn a stacked photograph b the same aedeagus, SEM image c detail of endophallic teeth d – f A. niger, GB, Hampshire, New Forest d stacked photograph e, f SEM images g – i A. wilsonae, SP, Areños g stacked photograph h SEM image, i detail of endophallic teeth j – l A. bameuli sp. nov., holotype j stacked photograph k SEM image l detail of endophallic teeth m – o A. krelli sp. nov., holotype m stacked photograph n SEM image o detail of endophallic teeth p – r A. bellumgerens sp. nov., paratype p stacked photograph q SEM image r detail of endophallic teeth s, t A. bellumgerens sp. nov., holotype s stacked photograph t SEM image of endophallus u A. muscorum, HU, Hortobágyi, stacked photograph v, w A. felix sp. nov., holotype, SEM images v aedeagus w detail of endophallic teeth. Black Scale bar: 0.5 mm for aedeagi and endophallus, white SEM scale on black background: 50 μm for details of endophallic teeth." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097492> ;
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dc:description "Figure 25. Aedeagal encasements (9 th abdominal sternite) a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. felix sp. nov., IT, Campo Felice d A. alberti sp. nov., TU, Ovit Dağı e A. bellumgerens sp. nov., IT, Sicily, Piano Battaglia f A. bameuli sp. nov., FR, Corsica, Haute-Corse g A. wilsonae, SP, Cantabria, Corconte h A. kraatzi, RU, Chita Reg. i A. plagiatus, UK, Norfolk, Hunstanton j A. plagiatus, CN, Beijing k A. rodrigoi sp. nov., SP, Madrid, Aranjuez l A. chellala sp. nov., AG, Chellala, paratype m A. rutilipennis sp. nov., CY, Konakli, paratype n A. discoides sp. nov., paratype. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097496> ;
a fabio:Figure .
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dc:description "Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097499> ;
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dc:description "Figure 32. Mitotic chromosomes arranged as karyotypes a – e A. felix sp. nov. IT, Abruzzo. Campo Felice a, b holotype ♂ a plain b the same nucleus C-banded c holotype ♂ C-banded, autosome 8 polymorphic, 9 subacrocentric d paratype ♂, C-banded, autosomes 8 and 9 metacentric e paratype ♀ f – i A. bellumgerens sp. nov., paratypes, IT, Sicily f, g A. bellumgerens sp. nov., ♂, paratype f plain g C-banded h, i A. bellumgerens sp. nov., ♀, paratype h plain i C-banded. Scale bar: 5 μm" ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097519> ;
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dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097509> ;
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dc:description "Figure 28. Habitat pictures a A. plagiatus, RU, Pavlosk Park b A. rodrigoi sp. nov., SP, Toledo, Quero c A. p. sinoplagiatus, CN, Tibet, Ganca d A. kraatzi, RU, Astrakhan oblast’ e A. alberti sp. nov., TU, Ovit Dağı (photograph Copyright Yunus Yasuv) f A. felix sp. nov., IT, Campo Felice." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097505> ;
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dwc:collectionCode "NHMUK" ;
dwc:eventDate "2009-06-01" ;
dwc:locality "Abruzzo" ;
dwc:recordedBy "R. B. Angus. Chromosome & R. B. Angus" ;
dwc:typeStatus "holotype" ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:title "Aphodius (Liothorax) rutilipennis Baudi di Selve 1870" ;
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bibo:volume "1207" ;
dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:date "2024" ;
dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
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dwc:authority "Baudi di Selve, 1870" ;
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dc:description "Figure 2. a – k whole beetles a A. plagiatus, CZ, Moravia b A. plagiatus, MG, Mongolia c A. jakutorum, holotype, RU, East Siberia d L. rodrigoi sp. nov., holotype SP, Aranjuez e A. rutilipennis (A. ressli, holotype) TR, Hatay, Iskenderun f A. rutilipennis, large ♀, CY, Akrotiri g A. rutilipennis (A. cypricola, holotype) CY h A. chellala sp. nov., holotype, AG, Chellala i A. rusakovi, ♂ paratype, RU, Orenburg j A. discoides, neotype (A. bytinskisalzi, holotype) IS, Kuneitra k A. discoides, black form from TR, Karakurt. Scale bar: 1 mm." ;
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dc:description "Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." ;
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dc:description "Figure 7. a – k epipharynxes a – f A. plagiatus plagiatus a, b GB, Hunstanton, Norfolk c CZ, Moravia, Dobré Pole d RU, Pavlovsk, St Petersburg e A. p. sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai f A. p. sinoplagiatus?, CH, “ Pekin, coll. Fry ” g A. rodrigoi sp. nov., holotype, SP h A. discoides, TR i A. rusakovi, paratype, RU, Orenburg j A. rutilipennis, CY, Akrotiri k A. chellala, sp. nov., paratype, AG, Chellala. Scale bar: 0.5 mm." ;
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dc:description "Figure 13. Aphodius plagiatus group, pronotal bases 1 a, b A. plagiatus a GB, Norfolk b MG c A. rutilipennis, CY d A. cf. rutilipennis, GR, Thessaloniki e A. chellala sp. nov., paratype, AG." ;
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dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
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dc:description "Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." ;
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dc:description "Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." ;
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dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:title "Aphodius (Liothorax) kraatzi Harold 1868" ;
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dc:creator "Angus, Robert B.; Maté, Jason F.; Angus, Elizabeth M.; Král, David" ;
dc:date "2024" ;
dc:title "Towards a revision of the Palaearctic species of Aphodius Hellwig, 1798, subgenus Liothorax Motschulsky, 1860 (Coleoptera, Scarabaeidae, Aphodiinae)" ;
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dc:description "Figure 3. a – i whole beetles a, b A. kraatzi a AF, Kabul, ♀ b KZ, Kyzil Kum c, d A. isikdagensis, TR, Ḉamildere c ♂ paratype d ♀ paratype e – g A. alberti sp. nov., TR. Rize, Ovitdagi e holotype ♂ f paratype ♂ g paratype ♀ h A. alberti?, ♂, AR, Armenia i A. isikdagensis?, ♂, TR, Artvin. Scale bar: 1 mm." ;
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dc:description "Figure 5. a – p A. plagiatus group, heads and pronota a – c A. plagiatus plagiatus a GB, Norfolk b CZ, Moravia, Dobre Pole c RU, St Petersburg d A. p. sinoplagiatus ssp. nov., paratype, CH, Qinghai, Gangca e – g A. discoides e neotype IS Kuneitra f TR, Kyzil Dag g Turkey Muş h A. discoides, black ♂, TR, Karakurt i – k A. rutilipennis i A. ressli, holotype, TR, Hatay j CY, Akrotiri k A. cypricola, holotype, CY l A. chellala sp. nov., holotype, AG, Chellala m A. rusakovi, paratype ♂, RU, Orenburg n A. ballerioi sp. nov., paratype ♂, TR, Rize o, p L. kraatzi o RU, Astrakhan p SR. Scale bar: 1 mm." ;
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dc:description "Figure 8. a – k epipharynxes a, b A. kraatzi, SV c, d A. isikdagensis paratypes, TR, Ḉamildere e, f A. felix sp. nov., paratypes g, h A. bameuli paratypes i A. alberti?, AR j A. krelli sp. nov., paratype k, l A. alberti sp. nov., paratypes. Scale bar: 0.5 mm" ;
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dc:description "Figure 14. Aphodius plagiatus group, pronotal bases 2 a A. discoides, TR b A. kraatzi, SK c L. rusakovi, paratype, RU, Volgograd oblast’." ;
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dc:description "Figure 17. Elytral sculpture of interstices 2 or 3 a – z stacked photographs a A. plagiatus plagiatus, CZ, Moravia b, c A. p. sinoplagiatus ssp. nov., paratypes, CH b Gangca, Qinghai c Gansu d A. rutilipennis (A. ressli, holotype), TR, Hatay e A. chellala sp. nov., holotype f A. rusakovi, paratype, RU, Orenburg g, h A. discoides g neotype, IS, Kuneitra h TR, Kizildag i A. discoides, black specimen, TR, Karakurt j A. kraatzi, SK k A. isikdagensis, paratype, TR, Ḉamildere l, m A. niger, Sweden, Södermanland, Tullgarn n – q A. muscorum, HU n holotype, Vörös-tó o Hortobágyi p, q Kisjúszállás r, s A. niger? r CZ, Hradec Králové s CZ, České Budějovice t A. muscorum? SK, Bol u A. bellumgerens sp. nov., paratype, IT, Sicily v A. felix sp. nov., paratype, IT w A. wilsonae, paratype, SP x A. bameuli sp. nov., paratype, FR, Corsica y A. krelli sp. nov., paratype, IT, Sardinia z A. alberti sp. nov., paratype, TR, Rize, Ovitdag. a’ – c ’ A. plagiatus, at higher magnification a ’, c ’ stacked photographs a ”, c ” SEM images a ’, a ” A. p. sinoplagiatus ssp. nov., CH, Gansu c ”, c ’ A. p. plagiatus, CZ, Moravia. Scale bars: 0.1 mm." ;
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dc:description "Figure 18. Legs a – h A. plagiatus group, stacked photographs mid tarsi and apical portions of tibiae a A. plagiatus sinoplagiatus ssp. nov., paratype, CH, Gangca, Qinghai b A. rodrigoi sp. nov. holotype, SP, Aranjuez c A. rutilipennis (A. ressli, holotype), TR, Hatay d A. chellala sp. nov., holotype, AG, Chellala e A. discoides, neotype, IS, Keneitra f, g A. kraatzi f SV g RU, Karasuk, W. Siberia h A. rusakovi, paratype ♂, RU, Orenburg i – t A. niger group, mid tarsi, and tibial apices i – o SEM images p – t stacked photographs i A. niger, Sweden, Södermanland, Tullgarn j A. muscorum, HU, Hortobágyi k, l A. bameuli sp. nov., paratypes, FR, Corsica m, n A. krelli sp. nov., paratypes, IT, Sardinia o A. wilsonae, paratype SP p, q A. felix sp. nov, IT, Abruzzo, Campo Felice p paratype q holotype r A. bellumgerens sp. nov., paratype, IT, Sicily s A. isikdagensis, paratype, TR, Ḉamildere t A. alberti sp. nov., holotype, TR, Rize, Ovitdag u, v fore tibiae and tarsi of ♂♂ to show the spur. u A. bameuli sp. nov., paratype, FR, Corsica v A. rusakovi, paratype, RU, Orenburg w, x apex of hind tibia to show the short spines w A. muscorum, HU x A. wilsonae, paratype, SP. Scale bar: 1 mm." ;
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dc:description "Figure 20. Metaventrites, A. plagiatus group a A. plagiatus plagiatus, GB, Studland, ♂ b A. p. sinoplagiatus ssp. nov., paratype, CH, Gansu, ♀ c, d A. rutilipennis, CY c ♂ d ♀ e, f A. chellala sp. nov., paratypes, AG e ♂ f ♀ g, h A. kraatzi, RU, Orenburg, ♂ i A. kraatzi, SK, ♀ j A. rodrigoi sp. nov., holotype, ♂, SP, Provinicia de Madrid, Aranjuez k, l A. discoides k neotype ♂, IS, Kuneitra l TR, ♀ m, n A. rusakovi m paratype, ♂, RU, Orenburg n ♀ RU, Volgograd oblast’. Scale bar: 0.5 mm." ;
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dc:description "Figure 22. Aphodius plagiatus group aedeagi a – c A. plagiatus plagiatus, GB, Studland, Dorset, aedeagus with extruded endophallus a, b SEM images a whole aedeagus b endophallus in more detail c stacked photograph of endophallus d – g A. discoides, neotype (Aphodius bytinskisalzi, holotype, IS, Kuneitra) d, e, e ’ SEM images f, g stacked photographs h – n cleared preparations mounted on slides h A. p. plagiatus, GB, Norfolk i A. p. sinoplagiatus? “ China, Pekin ”, Fry collection i ’ teeth on neck of endophallus in the region marked with an arrow in i j, k A. plagiatus sinoplagiatus ssp. nov., paratypes j CH, Gangca k CH, Tsaidam l Aphodius jakutorum Balthasar, lectotype, RU, East Siberia m A. rodrigoi sp. nov., holotype, SP, Aranjuez, dissected aedeagus m ’ endophallus m ” endophallic teeth in area indicated by arrow in m ’ at higher magnification n A. discoides, TR, Karacadag n ’ teeth on neck of endophallus, in area indicated by arrow in n at higher magnification o A. discoides, black specimen from TR, Karakurt, o ’ teeth on neck of endophallus, in area indicated by arrow in o at higher magnification p, p ’ A. rusakovi p paratype, RU, Orenburg p ’ RU Volgograd obl. Fastov, paramere, lateral q – v SEM images and stacked photographs q, r A. rutilipennis, CY, with endophallus inflated q SEM image q ’ (bottom right of plate) dorsal surface of endophallus in area indicated by arrow in q at higher magnification, to show the hair-like setae r stacked photograph of the same specimen s, s ’ A. chellala sp. nov., paratype with endophallus extruded s stacked photograph s ’, s ”, s ” ’ SEMs of the same specimen s ”, s ” ’ at higher magnifications t, u L. chellala sp. nov., holotype with the endophallus extruded t stacked photograph u SEM image v – z photographs of cleared specimens mounted in DMHF on slides v, w A. rutilipennis v CY w TR w ’ teeth on neck of endophallus, in area indicated by arrow in w at higher magnification x A. chellala sp. nov., paratype, AG, Taguin (La Smala) x ’ teeth on neck of endophallus, in area indicated by arrow in x at higher magnification y, z A. kraatzi y KZ, Syr Darya z AF, Kabul z ’ teeth on neck of endophallus, in area indicated by arrow in z at higher magnification Scale bar 1: 1 mm (a, d, f – t, v – z); 0.4 mm (n ’, o ’, s ”, w ’, x ’, z ’); 0.17 mm (e ’, m ”, q’ s ” ’). Scale bar 2: 1 mm (b, c, e, g and s ”)." ;
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<http://dx.doi.org/10.3897/zookeys.1207.117225.figure25>
dc:description "Figure 25. Aedeagal encasements (9 th abdominal sternite) a A. niger, GB, Hampshire, New Forest b A. muscorum, HU, Hortobagyi c A. felix sp. nov., IT, Campo Felice d A. alberti sp. nov., TU, Ovit Dağı e A. bellumgerens sp. nov., IT, Sicily, Piano Battaglia f A. bameuli sp. nov., FR, Corsica, Haute-Corse g A. wilsonae, SP, Cantabria, Corconte h A. kraatzi, RU, Chita Reg. i A. plagiatus, UK, Norfolk, Hunstanton j A. plagiatus, CN, Beijing k A. rodrigoi sp. nov., SP, Madrid, Aranjuez l A. chellala sp. nov., AG, Chellala, paratype m A. rutilipennis sp. nov., CY, Konakli, paratype n A. discoides sp. nov., paratype. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097496> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure26>
dc:description "Figure 26. Spermathecae a – e A. plagiatus plagiatus a CZ, Dobre Pole b GB, Norfolk c KZ, Karaganda d, e RU, Transbaikal, Ulan Ude f, g A. p. sinoplagiatus, CN f Gansu g holotype, CN, Qinghai h, i A. discoides, TR h Kizil dag, i Muʂ j A. rutilipennis CY, Limassol distr k A. rutilipennis?, GR Thesaloniki l A. chellala, paratype, AG, Chellala m, n A. rusakovi m paratype, RU, wet but not covered n KZ, Syr-Darja o – s A. kraatzi o, p SK q RU, Astrakhan r RU, Lenkoran s AF, Kabul t A. niger, GB, Hampshire, New Forest u A. muscorum HU, Hortobágyi v A felix, paratype, IT w, x A. bameuli, paratypes, FR y A. krelli, sp. nov., IT z A. bellumgerens sp. nov., paratype, IT a ’, b ’ A. alberti sp. nov., TR paratypes c ’ A. isikdagensis, paratype, TR Ḉamildere d ’, e ’ A. wilsonae, SP d ’ paratype, SP, Madrid e ’ SP, Soria. Scale bar: 0.5 mm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097499> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure30>
dc:description "Figure 30. Mitotic chromosomes, arranged as karyotypes a, b A. plagiatus plagiatus, ♂, GB, Dorset, Studland a plain (Giemsa stained) b the same nucleus C-banded c A. plagiatus sinoplagiatus, ♂, CH, Qinghai, Gangca, plain d, e A. rutilipennis, ♂, CY, Zakaki Marshes d plain e C-banded f – h A. kraatzi ♀, SK f, g nucleus without B-chromosomes f plain g C-banded h, i nucleus with 1 B-chromosome h plain i C-banded. Scale bar: 5 μm." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097512> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure29>
dc:description "Figure 29. a distribution of niger - group taxa in Western and Central Europe b distribution of Liothorax species in Central and Eastern Palearctic c distribution of plagiatus - group species in Central and Eastern Palearctic. The record of isikdagensis with the question mark indicates a doubtful specimen from Artvin Çam, TR (Ziani Coll)." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097509> ;
a fabio:Figure .
<http://dx.doi.org/10.3897/zookeys.1207.117225.figure28>
dc:description "Figure 28. Habitat pictures a A. plagiatus, RU, Pavlosk Park b A. rodrigoi sp. nov., SP, Toledo, Quero c A. p. sinoplagiatus, CN, Tibet, Ganca d A. kraatzi, RU, Astrakhan oblast’ e A. alberti sp. nov., TU, Ovit Dağı (photograph Copyright Yunus Yasuv) f A. felix sp. nov., IT, Campo Felice." ;
fabio:hasRepresentation <https://binary.pensoft.net/fig/1097505> ;
a fabio:Figure .