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dc: creator " C, Lees David " ;
dc: title " Heteropsis avaratra Lees & Kremen, sp. nov. " ;
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bibo: endPage " 97 " ;
bibo: issue " 1 " ;
bibo: journal " Zootaxa " ;
bibo: startPage " 1 " ;
bibo: volume " 4118 " ;
dc: creator " C, Lees David " ;
dc: date " 2016 " ;
dc: title " Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision " ;
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# Warning: Using document metadata for authority year ;
dwc: authority " Lees & Kremen " ;
dwc: authorityName " Lees & Kremen " ;
dwc: class " Insecta " ;
dwc: family " Curculionidae " ;
dwc: genus " Heteropsis " ;
dwc: kingdom " Animalia " ;
dwc: order " Coleoptera " ;
dwc: phylum " Arthropoda " ;
dwc: rank " species " ;
dwc: scientificNameAuthorship " Lees & Kremen, 2016 " ;
dwc: species " avaratra " ;
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dwc: class " Insecta " ;
dwc: family " Curculionidae " ;
dwc: genus " Heteropsis " ;
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dc: description " FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees & Kremen, sp. nov. (Montagne d’ Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’ Ambre, Les Roussettes; DCL-DB- 3385, MNHN). " ;
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dc: description " FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees & Kremen, sp. nov. (Montagne d’ Ambre). No wing preparation was available for Ht. sogai. " ;
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dc: description " FIGURE 16. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture (photos: D. C. Lees). A: HT ♂ of Heteropsis sogai, sp. nov. (Tsaratanana; DCL-DB- 3386). B: PT ♀ of Ht. sogai (Tsaratanana; DCL-DB- 3387). Scale bar: 30 mm. C: ♀ of Ht. kremenae Lees, sp. nov. (Vohitaly, Makira; 30 / 12 / 2002). D: ♂ of Ht. pauper (Vohitaly, Makira; 30 / 12 / 2002). E: ♀ of Ht. subsimilis (Ambohitantely; 6 / 12 / 2006); F: ♀ of Ht. avaratra Lees & Kremen, sp. nov. (Anjanaharibe Sud; 23 / 11 / 2014). " ;
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dc: description " FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees & Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3. " ;
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dc: description " FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees & Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm. " ;
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dc: description " FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees & Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’ ). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’ , BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades. " ;
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dc: description " FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees & Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale). " ;
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dc: description " FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees & Kremen, sp. nov. Light blue: Ht. hazovola Lees & Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees & Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees & Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees & Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees & Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees & Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui & Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees & Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km. " ;
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