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dc: creator " Duarte, Paulo R. M.; Grossi, Paschoal C.; Dupuis, Fabien " ;
dc: title " Bothynus spinophallicus Duarte & Grossi & Dupuis 2022, new combination " ;
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bibo: endPage " 66 " ;
bibo: issue " 1 " ;
bibo: journal " Zootaxa " ;
bibo: pubDate " 2022-01-25 " ;
bibo: startPage " 49 " ;
bibo: volume " 5093 " ;
dc: creator " Duarte, Paulo R. M.; Grossi, Paschoal C.; Dupuis, Fabien " ;
dc: date " 2022 " ;
dc: title " A taxonomic revision of the Bothynus villiersi Endrödi, 1968 species group (Coleoptera: Scarabaeidae: Dynastinae) " ;
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dwc: authority " (Ratcliffe & Cave, 2021) " ;
dwc: authorityName " Duarte & Grossi & Dupuis " ;
dwc: authorityYear " 2022 " ;
dwc: baseAuthorityName " Ratcliffe & Cave " ;
dwc: baseAuthorityYear " 2021 " ;
dwc: class " Insecta " ;
dwc: family " Dynastidae " ;
dwc: genus " Bothynus " ;
dwc: kingdom " Animalia " ;
dwc: order " Coleoptera " ;
dwc: phylum " Arthropoda " ;
dwc: rank " species " ;
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dwc: scientificNameAuthorship " (Ratcliffe & Cave, 2021) " ;
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dwc: species " spinophallicus " ;
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dwc: authority " Ratcliffe & Cave 2021: 45 " ;
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dwc: class " Insecta " ;
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dwc: authority " Endrodi, 1968 " ;
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dwc: authority " Endrodi, 1968 " ;
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dwc: authority " Martinez, 1983 " ;
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dwc: authority " Duarte & Grossi & Dupuis, 2022 " ;
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dwc: class " Insecta " ;
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dwc: authority " Hope, 1837 " ;
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dwc: authority " , Ratcliffe and Cave " ;
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dc: description " FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021). " ;
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dc: description " FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype). " ;
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dc: description " FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A– B, Bothynus villiersi Endrödi, 1968; C– D, Bothynus horridus Endrödi, 1968; E– F, Bothynus alvarengai Martínez, 1983; G– H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B. " ;
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dc: description " FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record. " ;
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dwc: collectionCode " UNSM " ;
dwc: locality " Paraguay " ;
dwc: recordedBy " R. C. Aguilar & Aguilar J. " ;
dwc: typeStatus " holotype " ;
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dwc: collectionCode " CERPE " ;
dwc: eventDate " 2011-05 " ;
dwc: locality " Parque Estadual do Cerrado " ;
dwc: recordedBy " C. S. Costa & Equipe " ;
trt: gbifOccurrenceId " 3455072345 " ;
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dwc: locality " Capiibary " ;
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