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dc: creator " Huber, Bernhard A.; Pavlek, Martina; Komnenov, Marjan " ;
dc: title " Stygopholcus photophilus Senglet 1971 " ;
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bibo: endPage " 60 " ;
bibo: journal " European Journal of Taxonomy " ;
bibo: pubDate " 2021-06-11 " ;
bibo: startPage " 1 " ;
bibo: volume " 752 " ;
dc: creator " Huber, Bernhard A.; Pavlek, Martina; Komnenov, Marjan " ;
dc: date " 2021 " ;
dc: title " Revision of the spider genus Stygopholcus (Araneae, Pholcidae), endemic to the Balkan Peninsula " ;
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dwc: authority " Senglet, 1971 " ;
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dwc: authorityYear " 1971 " ;
dwc: class " Arachnida " ;
dwc: family " Pholcidae " ;
dwc: genus " Stygopholcus " ;
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dwc: order " Araneae " ;
dwc: phylum " Arthropoda " ;
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dwc: species " photophilus " ;
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dc: description " Figs 1– 5. Stygopholcus Kratochvíl, 1932; live specimens. 1. S. absoloni (Kulczyński, 1914); male from Vjetrenica, Bosnia and Herzegovina. 2. S. skotophilus Kratochvíl, 1940; male from Pavlova pećina, Bosnia and Herzegovina. 3. S. montenegrinus Kratochvíl, 1940; male and female with egg-sac from Studenačka pećina, Montenegro. 4– 5. S. photophilus Senglet, 1971; male and female from Episkopi, Crete. Photos: BAH. " ;
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dc: description " Figs 6– 11. Stygopholcus Kratochvíl, 1932; left male pedipalps, prolateral, dorsal, and retrolateral views. 6– 8. S. skotophilus Kratochvíl, 1940; from Ilijina pećina, Bosnia and Herzegovina (ZFMK (Ar 22198)). 9– 11. S. photophilus Senglet, 1971; from Episkopi, Crete (ZFMK (Ar 22205)).Abbreviations: b = genital bulb; dp = dorsal process (of genital bulb); p = procursus; rp = retrolateral process (of genital bulb). Arrows point at joints shifted toward prolateral (femur-patella) and retrolateral (tibia-tarsus). Scale bars: 1 mm. " ;
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dc: description " Figs 145– 152. Stygopholcus Kratochvíl, 1932; cleared female genitalia, ventral and dorsal views. 145– 146. S. absoloni (Kulczyński, 1914); from Zavala, Bosnia and Herzegovina (MHNG). 147– 148. S. skotophilus Kratochvíl, 1940; from Pećina u Ivici, Montenegro ((ZFMK Ar 22196)). 149– 150. S. montenegrinus Kratochvíl, 1940; from Golubinja pećina, Montenegro (CMK). 151– 152. S. photophilus Senglet, 1971; from Choumerion/Pérama, Crete (MHNG). " ;
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dc: description " Figs 153– 156. Stygopholcus Kratochvíl, 1932; cleared female genitalia in dorsal views; dorsal sclerotized arcs tilted towards the back. 153. S. absoloni (Kulczyński, 1914); from Balićeva špilja, Croatia (ZFMK (Ar 22195)). 154. S. skotophilus Kratochvíl, 1940; from Pećina u Ivici, Montenegro (ZFMK (Ar 22196)). 155. S. montenegrinus Kratochvíl, 1940; from Studenačka pećina, Montenegro (ZFMK (Ar 22201)). 156. S. photophilus Senglet, 1971; from Choumerion/Pérama, Crete (MHNG). Abbreviations: da = dorsal arc; e = epigynal plate; v = ʻ valve’ ; va = ventral arc. " ;
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dc: description " Figs 123– 128. Stygopholcus photophilus Senglet, 1971; epigyna, ventral views, and details of male femora 2 and 3, dorsal views. 123. Thessaly, Omolio, Tempi Valley (MHNG). 124. Corfu, Loutses (MHNG). 125. Peloponnese, Agios Dimitrios/Lefktro (MHNG); arrows point at internal sclerites. 126– 128. Crete, Episkopi (ZFMK (Ar 22205)). Scale bars: 123– 126 = 0.5 mm; 127– 128 = 0.2 mm. " ;
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dc: description " Figs 129– 136. Stygopholcus photophilus Senglet, 1971; male and female from Choumerion/Pérama, Crete (MHNG). 129. Male clypeus, frontal view; arrow points at modified median area. 130. Male chelicerae, frontal view. 131. Modified area medially on male chelicerae. 132. Modified hairs frontally on male chelicerae. 133. Stridulatory files on male chelicerae. 134. Absence of stridulatory files on female chelicerae. 135. Male ALS. 136. Female anterior lateral spinneret (ALS) and posterior median spinneret (PMS). Scale bars: 129– 130 = 100 µm; 131, 134– 136 = 20 µm; 132– 133 = 10 µm. " ;
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dc: description " Figs 161– 166. Procursus length, number of modified hairs on each male chelicera, epigynum width/ length, epigynum measure F (cf. Fig. 21), and relationships between tibia 1 and tibia 2 lengths in males and females. Measures/counts sorted from smallest to biggest for each species. Abbreviations: abs. = S. absoloni; sko. = S. skotophilus; mon. = S. montenegrinus; pho. = S. photophilus. " ;
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dc: description " Figs 113– 118. Stygopholcus photophilus Senglet, 1971; male from Episkopi, Crete (ZFMK (Ar 22205)). 113– 114. Left tarsus and procursus, prolateral and retrolateral views. 115– 118. Left genital bulb, retrolateral, ventral, prolateral-distal, and dorso-distal views; arrow points at weakly sclerotized process. Abbreviations: bs = basal sclerite; dp = dorsal process; ds = dorsal sclerite; rp = retrolateral process; vs = ventral sclerite. Scale bars: 0.5 mm. " ;
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dc: description " Figs 137– 144. Stygopholcus photophilus Senglet, 1971; male and female from Choumerion/Pérama, Crete (MHNG). 137– 138. Tip of left procursus, retrolateral (slightly ventral) and ventral views. 139. Detail of previous figure. 140. Dorsal bulbal process and transparent bulbal process (arrow); note hair-like structures on dorsal bulbal process. 141. Female palpal tarsal organ. 142. Tarsal organ on female tarsus 4. 143. Male gonopore and epiandrous spigots. 144. Anterior part of epigynum showing strong ridges; arrows point at pair of anterior bulges. Abbreviations: ds = dorsal sclerite; vs = ventral sclerite. Scale bars: 137– 138, 140, 143– 144 = 100 µm; 139, 141– 142 = 10 µm. " ;
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dc: description " Figs 119– 120. Stygopholcus photophilus Senglet, 1971; male from Episkopi, Crete (ZFMK (Ar 22205)), female from Choumerion/Pérama, Crete (MHNG). 119– 120. Male chelicerae, frontal and lateral views. 121– 122. Cleared female genitalia, ventral and dorsal views. Scale bars: 0.5 mm. " ;
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dc: description " Fig. 167. Known distribution of Stygopholcus Kratochvíl, 1932. For the ‘ northern clade’ , see Fig. 168. Abbreviations: Alb = Albania; BaH = Bosnia and Herzegovina; Cro = Croatia; Gre = Greece; Mon = Montenegro. " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1976-05-14 " ;
dwc: locality " Thessaly " ;
dwc: recordedBy " B. Hauser " ;
dwc: verbatimElevation " 600 " ;
dwc: verbatimLatitude " 39.862 " ;
dwc: verbatimLongitude " 22.572 " ;
trt: gbifOccurrenceId " 3302592558 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1978-04-10 " ;
dwc: locality " Kri " ;
dwc: recordedBy " B. Hauser " ;
dwc: verbatimElevation " 600 " ;
dwc: verbatimLatitude " 39.862 " ;
dwc: verbatimLongitude " 22.572 " ;
trt: gbifOccurrenceId " 3302592548 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1978-04-11 " ;
dwc: locality " Tsari Tripa cave " ;
dwc: recordedBy " B. Hauser " ;
dwc: verbatimLatitude " 39.821 " ;
dwc: verbatimLongitude " 22.641 " ;
trt: gbifOccurrenceId " 3302592600 " ;
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dwc: catalogNumber " Ar 22202 " ;
dwc: collectionCode " ZFMK " ;
dwc: eventDate " 1995-09-22 " ;
dwc: locality " Stomio " ;
dwc: recordedBy " K. Thaler & B. Knoflach " ;
dwc: verbatimLatitude " 39.86 " ;
dwc: verbatimLongitude " 22.72 " ;
trt: gbifOccurrenceId " 3302592342 " ;
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dwc: catalogNumber " Ar 22203 " ;
dwc: collectionCode " ZFMK " ;
dwc: eventDate " 1969-05-04 " ;
dwc: locality " Dafni " ;
dwc: recordedBy " Ausobsky " ;
dwc: verbatimElevation " 10 " ;
dwc: verbatimLatitude " 40.213 " ;
dwc: verbatimLongitude " 24.223 " ;
trt: gbifOccurrenceId " 3302592462 " ;
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dwc: collectionCode " NMNHS " ;
dwc: eventDate " 2012-09-08 " ;
dwc: locality " Lost Glasses Cave " ;
dwc: recordedBy " A. Zhalov " ;
dwc: verbatimLatitude " 40.14 " ;
dwc: verbatimLongitude " 24.3 " ;
trt: gbifOccurrenceId " 3302592595 " ;
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dwc: collectionCode " SMF " ;
dwc: eventDate " 2009-08-22 " ;
dwc: locality " Sithonia " ;
dwc: recordedBy " S. Huber & A. Schonhofer & Eastern " ;
dwc: verbatimElevation " 20 " ;
dwc: verbatimLatitude " 40.1702 " ;
dwc: verbatimLongitude " 23.8545 " ;
trt: gbifOccurrenceId " 3302592382 " ;
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dwc: collectionCode " NMNHS " ;
dwc: eventDate " 1982-12-28 " ;
dwc: locality " Drakotrypa cave " ;
dwc: recordedBy " P. Beron & S. Andreev " ;
dwc: verbatimLatitude " 40.732 " ;
dwc: verbatimLongitude " 24.732 " ;
trt: gbifOccurrenceId " 3302592388 " ;
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dwc: collectionCode " NMNHS " ;
dwc: eventDate " 1982-12-28 " ;
dwc: locality " Aghia Helleni cave " ;
dwc: recordedBy " P. Beron & S. Andreev " ;
dwc: verbatimLatitude " 41.018 " ;
dwc: verbatimLongitude " 24.394 " ;
trt: gbifOccurrenceId " 3302592490 " ;
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dwc: collectionCode " SMF " ;
dwc: eventDate " 2009-08-14 " ;
dwc: locality " Monodendri " ;
dwc: recordedBy " S. Huber & A. Schonhofer " ;
dwc: verbatimElevation " 1000 " ;
dwc: verbatimLatitude " 39.883 " ;
dwc: verbatimLongitude " 20.749 " ;
trt: gbifOccurrenceId " 3302592498 " ;
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dwc: catalogNumber " Ar 22204 " ;
dwc: collectionCode " ZFMK " ;
dwc: eventDate " 2006-09-29 " ;
dwc: locality " 12 km Wof Leonidion " ;
dwc: recordedBy " Ag. Taxiarchi & H. Pieper " ;
dwc: verbatimLatitude " 37.15 " ;
dwc: verbatimLongitude " 22.76 " ;
trt: gbifOccurrenceId " 3302592476 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1970-08-16 " ;
dwc: locality " Sparti " ;
dwc: recordedBy " A. Senglet " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 37.067 " ;
dwc: verbatimLongitude " 22.375 " ;
trt: gbifOccurrenceId " 3302592303 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1983-05-23 " ;
dwc: locality " Mystras " ;
dwc: recordedBy " A. Senglet " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 37.067 " ;
dwc: verbatimLongitude " 22.375 " ;
trt: gbifOccurrenceId " 3302592480 " ;
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dwc: collectionCode " MCVR " ;
dwc: eventDate " 2016-12-05 " ;
dwc: locality " Spilaio Skreti " ;
dwc: recordedBy " C. Cenni " ;
dwc: typeStatus " paratype " ;
dwc: verbatimElevation " 490 " ;
dwc: verbatimLatitude " 37.1676 " ;
dwc: verbatimLongitude " 22.3052 " ;
trt: gbifOccurrenceId " 3302592477 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1970-08-20 " ;
dwc: locality " Lefktro " ;
dwc: recordedBy " A. Senglet " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 37.039 " ;
dwc: verbatimLongitude " 22.102 " ;
trt: gbifOccurrenceId " 3302592458 " ;
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dwc: catalogNumber " 1845 " ;
dwc: collectionCode " HISR " ;
dwc: eventDate " 1997-09-29 " ;
dwc: locality " Spilaio Kapsias " ;
dwc: recordedBy " K. Paragamian " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 37.623 " ;
dwc: verbatimLongitude " 22.354 " ;
trt: gbifOccurrenceId " 3302592365 " ;
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dwc: collectionCode " HISR " ;
dwc: eventDate " 1997-09-19 " ;
dwc: locality " Spilaio Dionysou " ;
dwc: recordedBy " K. Paragamian " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 37.1485 " ;
dwc: verbatimLongitude " 22.8185 " ;
trt: gbifOccurrenceId " 3302592596 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1970-07-01 " ;
dwc: locality " Perama " ;
dwc: recordedBy " A. Senglet " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 35.355 " ;
dwc: verbatimLongitude " 24.72 " ;
trt: gbifOccurrenceId " 3302592605 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1970-07-02 " ;
dwc: locality " Perama " ;
dwc: recordedBy " A. Senglet " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 35.333 " ;
dwc: verbatimLongitude " 24.74 " ;
trt: gbifOccurrenceId " 3302592431 " ;
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dwc: catalogNumber " 56590 " ;
dwc: collectionCode " SMF " ;
dwc: eventDate " 2006-05-17 " ;
dwc: locality " Archaia Eleftherna " ;
dwc: recordedBy " H. Eikamp & U Kluge " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 35.324 " ;
dwc: verbatimLongitude " 24.676 " ;
trt: gbifOccurrenceId " 3302592626 " ;
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dwc: catalogNumber " 1640 " ;
dwc: collectionCode " HISR " ;
dwc: eventDate " 1987-03-01 " ;
dwc: locality " Spilaio Skoteinospilios " ;
dwc: recordedBy " K. Paragamian " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 35.336 " ;
dwc: verbatimLongitude " 24.505 " ;
trt: gbifOccurrenceId " 3302592560 " ;
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dwc: collectionCode " NHMC " ;
dwc: eventDate " 2015-07-07 " ;
dwc: locality " Episkopi " ;
dwc: recordedBy " B. A. Huber " ;
dwc: typeStatus " paratype " ;
dwc: verbatimElevation " 230 " ;
dwc: verbatimLatitude " 35.3417 " ;
dwc: verbatimLongitude " 24.7656 " ;
trt: gbifOccurrenceId " 3302592478 " ;
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dwc: catalogNumber " Ar 22205 " ;
dwc: collectionCode " ZFMK " ;
dwc: eventDate " 2015-07-07 " ;
dwc: locality " Episkopi " ;
dwc: recordedBy " Ar " ;
dwc: typeStatus " paratype " ;
dwc: verbatimElevation " 230 " ;
dwc: verbatimLatitude " 35.3417 " ;
dwc: verbatimLongitude " 24.7656 " ;
trt: gbifOccurrenceId " 3302592447 " ;
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dwc: catalogNumber " Cre19 " ;
dwc: collectionCode " ZFMK " ;
dwc: eventDate " 2015-07-07 " ;
dwc: locality " Episkopi " ;
dwc: recordedBy " Cre " ;
dwc: typeStatus " paratype " ;
dwc: verbatimElevation " 230 " ;
dwc: verbatimLatitude " 35.3417 " ;
dwc: verbatimLongitude " 24.7656 " ;
trt: gbifOccurrenceId " 3302592319 " ;
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dwc: collectionCode " SMF " ;
dwc: eventDate " 2009-05-07 " ;
dwc: locality " Zaros " ;
dwc: recordedBy " K. Eckl & H. Eikamp " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 35.13 " ;
dwc: verbatimLongitude " 24.9 " ;
trt: gbifOccurrenceId " 3302592318 " ;
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dwc: catalogNumber " 2008 " ;
dwc: collectionCode " HISR " ;
dwc: eventDate " 2001-12-30 " ;
dwc: locality " Spilaio Agias Sofias " ;
dwc: recordedBy " K. Paragamian " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 35.417 " ;
dwc: verbatimLongitude " 23.683 " ;
trt: gbifOccurrenceId " 3302592406 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1999-10-10 " ;
dwc: locality " Spilaio Agias Sofias " ;
dwc: recordedBy " A. Senglet " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 35.417 " ;
dwc: verbatimLongitude " 23.683 " ;
trt: gbifOccurrenceId " 3302592586 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1972-08-31 " ;
dwc: locality " Istiaia " ;
dwc: recordedBy " A. Senglet " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 39.02 " ;
dwc: verbatimLongitude " 23.25 " ;
trt: gbifOccurrenceId " 3302592518 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1972-09-02 " ;
dwc: locality " Kato Steni " ;
dwc: recordedBy " A. Senglet " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 38.583 " ;
dwc: verbatimLongitude " 23.85 " ;
trt: gbifOccurrenceId " 3302592373 " ;
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dwc: collectionCode " SMF " ;
dwc: eventDate " 2006-03-27 " ;
dwc: locality " Lefkada " ;
dwc: recordedBy " A. Schonhofer " ;
dwc: typeStatus " paratype " ;
dwc: verbatimElevation " 200 " ;
dwc: verbatimLatitude " 38.7263 " ;
dwc: verbatimLongitude " 20.6858 " ;
trt: gbifOccurrenceId " 3302592540 " ;
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dwc: collectionCode " MMUE " ;
dwc: eventDate " 1987-05-28 " ;
dwc: locality " Lake Avithos " ;
dwc: recordedBy " J. Murphy " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 38.109 " ;
dwc: verbatimLongitude " 20.528 " ;
trt: gbifOccurrenceId " 3302592320 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1972-09-20 " ;
dwc: locality " Loutses " ;
dwc: recordedBy " A. Senglet " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 39.786 " ;
dwc: verbatimLongitude " 19.88 " ;
trt: gbifOccurrenceId " 3302592413 " ;
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dwc: collectionCode " SMF " ;
dwc: eventDate " 2006-03-22 " ;
dwc: locality " Peristerograva " ;
dwc: recordedBy " A. Schonhofer " ;
dwc: typeStatus " paratype " ;
dwc: verbatimElevation " 350 " ;
dwc: verbatimLatitude " 39.777 " ;
dwc: verbatimLongitude " 19.89 " ;
trt: gbifOccurrenceId " 3302592314 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1960-04-19 " ;
dwc: locality " Halbhohle am Nordabhang " ;
dwc: recordedBy " B. Hauser " ;
dwc: typeStatus " paratype " ;
dwc: verbatimElevation " 350 " ;
dwc: verbatimLatitude " 39.777 " ;
dwc: verbatimLongitude " 19.89 " ;
trt: gbifOccurrenceId " 3302592367 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1974-05-12 " ;
dwc: locality " Spilaio Katsampa " ;
dwc: recordedBy " B. Hauser " ;
dwc: typeStatus " paratype " ;
dwc: verbatimLatitude " 39.744 " ;
dwc: verbatimLongitude " 19.839 " ;
trt: gbifOccurrenceId " 3302592519 " ;
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dwc: collectionCode " NMNHS " ;
dwc: eventDate " 1995-05-12 " ;
dwc: locality " Permet " ;
dwc: recordedBy " B. Petrov & R. Stoev " ;
dwc: verbatimLatitude " 40.208 " ;
dwc: verbatimLongitude " 20.419 " ;
trt: gbifOccurrenceId " 3302592544 " ;
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dwc: collectionCode " MHNG " ;
dwc: eventDate " 1973-04-30 " ;
dwc: locality " Epirus " ;
dwc: recordedBy " V. Mahnert " ;
dwc: verbatimLatitude " 40.019 " ;
dwc: verbatimLongitude " 20.785 " ;
trt: gbifOccurrenceId " 3302592551 " ;
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dwc: eventDate " 2020-09-30 " ;
dwc: locality " Durres " ;
dwc: recordedBy " C. Deltshev " ;
trt: gbifOccurrenceId " 3302592593 " ;
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dwc: eventDate " 1993-05-26 " ;
dwc: locality " Durres " ;
dwc: recordedBy " P. Stoev & D. Zapryanova " ;
dwc: verbatimLatitude " 41.3 " ;
dwc: verbatimLongitude " 19.5 " ;
trt: gbifOccurrenceId " 3302592377 " ;
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dwc: eventDate " 1994-06-01 " ;
dwc: locality " Korce " ;
dwc: recordedBy " T. Ivanova " ;
dwc: verbatimLatitude " 40.15 " ;
dwc: verbatimLongitude " 20.6 " ;
trt: gbifOccurrenceId " 3302592612 " ;
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dwc: eventDate " 1994-06-01 " ;
dwc: locality " Vijoshe River " ;
dwc: recordedBy " G. Blagoev " ;
dwc: verbatimLatitude " 40.288 " ;
dwc: verbatimLongitude " 20.069 " ;
trt: gbifOccurrenceId " 3302592514 " ;
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dwc: collectionCode " SMF " ;
dwc: eventDate " 1926-05 " ;
dwc: locality " Crete " ;
dwc: recordedBy " Roewer " ;
trt: gbifOccurrenceId " 3302592459 " ;
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dwc: catalogNumber " Ar 10230 " ;
dwc: collectionCode " MNHN " ;
dwc: typeStatus " paratype " ;
trt: gbifOccurrenceId " 3302592457 " ;
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dwc: collectionCode " SMF " ;
dwc: eventDate " 1980-10-06 " ;
dwc: locality " Peloponnese " ;
dwc: recordedBy " Malicky " ;
dwc: typeStatus " paratype " ;
trt: gbifOccurrenceId " 3302592499 " ;
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