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dc: creator " Duarte, Paulo R. M.; Grossi, Paschoal C.; Dupuis, Fabien " ;
dc: title " Bothynus belemensis Duarte, Grossi, & Dupuis 2022, new species " ;
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bibo: endPage " 66 " ;
bibo: issue " 1 " ;
bibo: journal " Zootaxa " ;
bibo: pubDate " 2022-01-25 " ;
bibo: startPage " 49 " ;
bibo: volume " 5093 " ;
dc: creator " Duarte, Paulo R. M.; Grossi, Paschoal C.; Dupuis, Fabien " ;
dc: date " 2022 " ;
dc: title " A taxonomic revision of the Bothynus villiersi Endrödi, 1968 species group (Coleoptera: Scarabaeidae: Dynastinae) " ;
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dwc: authority " Duarte & Grossi & Dupuis, 2022 " , " Duarte, Grossi, & Dupuis " ;
dwc: authorityName " Duarte & Grossi & Dupuis " , " Duarte, Grossi, & Dupuis " ;
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dwc: authorityYear " 2022 " ;
dwc: class " Insecta " ;
dwc: family " Dynastidae " ;
dwc: genus " Bothynus " ;
dwc: kingdom " Animalia " ;
dwc: order " Coleoptera " ;
dwc: phylum " Arthropoda " ;
dwc: rank " species " ;
dwc: scientificNameAuthorship " Duarte, Grossi, & Dupuis, 2022 " ;
dwc: species " belemensis " ;
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dwc: authority " Endrodi, 1968 " ;
dwc: authorityName " Endrodi " ;
dwc: authorityYear " 1968 " ;
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dwc: class " Insecta " ;
dwc: family " Dynastidae " ;
dwc: genus " Bothynus " ;
dwc: kingdom " Animalia " ;
dwc: order " Coleoptera " ;
dwc: phylum " Arthropoda " ;
dwc: rank " species " ;
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dwc: scientificNameAuthorship " Endrodi, 1968 " ;
dwc: species " villiersi " ;
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dwc: authority " Endrodi, 1968 " ;
dwc: authorityName " Endrodi " ;
dwc: authorityYear " 1968 " ;
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dwc: class " Insecta " ;
dwc: family " Dynastidae " ;
dwc: genus " Bothynus " ;
dwc: kingdom " Animalia " ;
dwc: order " Coleoptera " ;
dwc: phylum " Arthropoda " ;
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dwc: rank " species " ;
dwc: scientificNameAuthorship " Endrodi, 1968 " ;
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dwc: authority " Martinez, 1983 " ;
dwc: authorityName " Martinez " ;
dwc: authorityYear " 1983 " ;
dwc: class " Insecta " ;
dwc: family " Dynastidae " ;
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dwc: order " Coleoptera " ;
dwc: phylum " Arthropoda " ;
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dwc: class " Insecta " ;
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dwc: genus " Bothynus " ;
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dwc: kingdom " Animalia " ;
dwc: order " Coleoptera " ;
dwc: phylum " Arthropoda " ;
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dwc: rank " species " ;
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dc: description " FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype). " ;
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dc: description " FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D. " ;
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dc: description " FIGURE 5. Spiculum gastrale: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. hs = hemisternite. " ;
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dc: description " FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A– – – – " ;
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dc: description " FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. " ;
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dc: description " FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate. " ;
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dc: description " FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record. " ;
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dwc: locality " Benedicto Nogueira " ;
dwc: typeStatus " holotype " ;
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